Physaria acutifolia |
Physaria rectipes |
|
---|---|---|
double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
straight bladderpod |
|
Habit | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | Perennials; (loose, spreading); caudex simple or branched; densely pubescent, trichomes (subsessile), 4–6-rayed, rays furcate or bifurcate, (moderately tuberculate over arms, less so or smooth over center). |
Stems | several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
few to several from base, ascending or prostrate, (arising laterally, also from within basal leaves, usually unbranched, rarely branched), 0.5–3(–6) dm. |
Basal leaves | (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
blade narrowly oblanceolate or broadly elliptic, 1–7(–12) cm, margins entire or shallowly toothed, sometimes repand, (inner blades usually flattened in age, surfaces often gray-green, scabrous). |
Cauline leaves | blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
(usually secund); blade spatulate or obovate, 1–2.5(–4.5) cm, margins entire or shallowly toothed, (flat or involute). |
Racemes | loose, (elongated in fruit). |
somewhat crowded (to moderately elongated in fruit, exceeding basal leaves). |
Flowers | sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
sepals broadly elliptic or oblong, 4–7.5(–9) mm, (median pair thickened apically, cucullate); petals cuneate or obovate, 7–10(–16) mm, (tapering gradually to broad claw). |
Fruiting pedicels | (divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
(often divaricate-spreading and straight, or horizontal and loosely sigmoid, sometimes slightly recurved), 5–15 mm. |
Fruits | (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
subglobose to ovoid or ellipsoid, sometimes compressed, (4–)5–7(–9) mm; valves sparsely pubescent, trichomes appressed or erect, sometimes sparsely pubescent inside; ovules (8–)12–16(–20) per ovary; style 2–7 mm. |
Seeds | (dark brown), flattened, (2–3 mm). |
somewhat flattened. |
2n | = 10, 16, 24. |
= 10 + 2, 18, 20, ± 40. |
Physaria acutifolia |
Physaria rectipes |
|
Phenology | Flowering May–Jun(-Jul). | Flowering May–Jul. |
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | Sandy soils, limey knolls, rocky hills, clay hillsides, dry ridges, weathered rocks, gravelly outwashes, stony slopes, pinyon-juniper woodlands |
Elevation | 1500-3500 m (4900-11500 ft) | 1500-2600 m (4900-8500 ft) |
Distribution |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
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AZ; CO; NM; UT
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Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
As here circumscribed, Physaria rectipes remains heterogeneous and may represent more than one taxon, even after the recent removal of P. pulvinata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 660. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | Lesquerella rectipes |
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | (Wooton & Standley) O’Kane & Al-Shehbaz: Novon 12: 327. (2002) |
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