Physaria acutifolia |
Physaria pycnantha |
|
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double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
mountain-view bladderpod |
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Habit | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | Perennials; caudex branched, (densely cespitose and forming hemispheric mounds); densely pubescent, trichomes 5-rayed, rays bifurcate near base, fused at base, (strongly tuberculate throughout). |
Stems | several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
few to several from base, erect, (usually exceeding basal leaves), 0.3–0.7 dm. |
Basal leaves | (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
blade linear-spatulate, 1.5–4 cm, (base narrowed gradually to petiole), margins entire. |
Cauline leaves | blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
blade spatulate, similar to basal. |
Racemes | loose, (elongated in fruit). |
crowded in distal 1/3, (4–10-flowered). |
Flowers | sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
sepals (pale yellow), oblong to elliptic, 3–4 mm, (median pair usually thickened apically, cucullate); petals (sometimes with slight tinge of orange basally), lingulate, 4–6 mm. |
Fruiting pedicels | (divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
(loosely to strongly sigmoid), 6–10 mm. |
Fruits | (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
ellipsoid, slightly inflated (somewhat latiseptate), 4–5 mm, (apex acute); valves pubescent, trichomes erect, appearing slightly shaggy; ovules 4–8 per ovary; styles 2.5–3 mm, (shorter than mature fruits). |
Seeds | (dark brown), flattened, (2–3 mm). |
± flattened, convex on outer side. |
2n | = 10, 16, 24. |
|
Physaria acutifolia |
Physaria pycnantha |
|
Phenology | Flowering May–Jun(-Jul). | Flowering late May–Jun(-Jul). |
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | Dry, windswept knolls of limestone gravel, with other cushion-forming plants |
Elevation | 1500-3500 m (4900-11500 ft) | 1600-2300 m (5200-7500 ft) |
Distribution |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
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ID; MT
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Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria pycnantha is morphologically similar to 56. P. nelsonii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 659. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | |
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | Grady & O’Kane: Novon 17: 188, fig. 5. (2007) |
Web links |