Physaria acutifolia |
Physaria pulvinata |
|
---|---|---|
double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
cushion bladderpod |
|
Habit | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | Perennials; caudex (buried), branched, (dense, forming hard mats); densely pubescent, trichomes (subsessile), 8–13-rayed, rays usually furcate, distinct, (umbonate, usually tuberculate, less so over umbo). |
Stems | several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
several (to several hundred) from base, erect, (each terminating in a tufted cluster of leaves), to 7 dm. |
Basal leaves | (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
(petiole not differentiated from blade); blade narrowly elliptic to narrowly linear-oblanceolate, (0.8–)1–1.5 cm, (base cuneate), margins entire. |
Cauline leaves | blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
similar to basal, blade sometimes linear, (apex acute). |
Racemes | loose, (elongated in fruit). |
dense, (often ± subumbellate, somewhat elongated in fruit). |
Flowers | sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
sepals narrowly elliptic, 2.5–3.5(–4) mm, (not keeled); petals narrowly spatulate, 4–7 mm. |
Fruiting pedicels | (divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
(strongly sigmoid), 5–10 mm. |
Fruits | (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
ellipsoid, compressed, 4–6 mm; valves densely pubescent, trichomes appressed; ovules 2 per ovary; style 2–3.5 mm. |
Seeds | (dark brown), flattened, (2–3 mm). |
flattened, (oval). |
2n | = 10, 16, 24. |
|
Physaria acutifolia |
Physaria pulvinata |
|
Phenology | Flowering May–Jun(-Jul). | Flowering late May–Jun. |
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | Gray, argillaceous shale outcrops with sagebrush and junipers |
Elevation | 1500-3500 m (4900-11500 ft) | 2300-2600 m (7500-8500 ft) |
Distribution |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
|
CO |
Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria pulvinata is known from an area surrounded by a pygmy forest of Utah juniper in Dolores and San Miguel Counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 659. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | |
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | O’Kane & Reveal: Brittonia 58: 74, fig. 1. (2006) |
Web links |