Physaria acutifolia |
Physaria pruinosa |
|
---|---|---|
double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
frosty bladderpod, Pagosa bladderpod |
|
Habit | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | Perennials; caudex simple or branched, (covered with persistent leaf bases); densely pubescent, trichomes (sessile or subsessile), 4–7-rayed, rays furcate or bifurcate, (tuberculate throughout). |
Stems | several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
simple or several from base, decumbent or erect, (unbranched), to 2 dm. |
Basal leaves | (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
(petiole sharply differentiated from blade, slender); blade suborbicular or obovate to rhombic, 4–8 cm, (base abruptly narrowed to petiole), margins entire, sinuate, or shallowly dentate, (abaxial surface densely pubescent, adaxial lightly pubescent). |
Cauline leaves | blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
(proximal petiolate, distal sessile); blade obovate to rhombic, 0.8–2.3 cm, margins entire or shallowly toothed. |
Racemes | loose, (elongated in fruit). |
dense, (somewhat elongated in fruit). |
Flowers | sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
sepals elliptic or oblong, ca. 6 mm, (lateral pair not saccate or subsaccate, cucullate, median pair thickened apically, cucullate); petals spatulate, ca. 9 mm, (claw expanded at base). |
Fruiting pedicels | (divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
(horizontal to ascending, sigmoid or slightly curved), 8–11 mm, (stout). |
Fruits | (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
(sessile or substipitate, often becoming copper-red in age), subglobose or ellipsoid, inflated, 6–9 mm, (firm, glossy); valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–8(–12) per ovary; style 3.5–7 mm. |
Seeds | (dark brown), flattened, (2–3 mm). |
somewhat flattened. |
2n | = 10, 16, 24. |
|
Physaria acutifolia |
Physaria pruinosa |
|
Phenology | Flowering May–Jun(-Jul). | Flowering May–Jun(-Aug). |
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | Mancos slate or shale, meadows, gentle slopes, edges of ponderosa pine stands |
Elevation | 1500-3500 m (4900-11500 ft) | 2100-2600 m (6900-8500 ft) |
Distribution |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
|
CO; NM |
Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. The one New Mexico population is near the border with Colorado, in Rio Arriba County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 658. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | Lesquerella pruinosa |
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | (Greene) O’Kane & Al-Shehbaz: Novon 12: 327. (2002) |
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