Physaria acutifolia |
Physaria pinetorum |
|
---|---|---|
double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
White Mountain bladderpod |
|
Habit | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | Perennials; caudex simple or branched; densely pubescent, trichomes (sessile or short-stalked), 6–8-rayed, rays furcate or bifurcate, (tuberculate, less so on outer layers). |
Stems | several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
simple or few from base, ascending to erect, (0.5–)1–2(–3.5) dm. |
Basal leaves | (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
(petiole tapering to blade); blade rhombic to elliptic and irregularly angular, sometimes spatulate to oblanceolate, 1.5–7.5(–10) cm, margins entire. |
Cauline leaves | blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
(not or loosely overlapping, petiolate or distal sessile); blade spatulate to oblanceolate, 1–4 cm, margins entire. |
Racemes | loose, (elongated in fruit). |
crowded, elongated. |
Flowers | sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
sepals ovate, oblong, or elliptic 4–7.5 mm, (median pair thickened apically, cucullate); petals spatulate or broadly cuneate, 6–13 mm, (claw slightly expanded at base). |
Fruiting pedicels | (divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
(ascending, curved or sigmoid), 6–12(–20) mm. |
Fruits | (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
(substipitate), globose or obovoid to ellipsoid, sometimes slightly obcompressed, 4–9 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4–24 per ovary; style (2–)4–7 mm. |
Seeds | (dark brown), flattened, (2–3 mm). |
flattened. |
2n | = 10, 16, 24. |
= 10. |
Physaria acutifolia |
Physaria pinetorum |
|
Phenology | Flowering May–Jun(-Jul). | Flowering Apr–Jul. |
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | Scrub oak, pinyon-juniper woodland, open ponderosa pine forests, these sometimes mixed with Douglas fir, white pine, white fir, Engelmann spruce, or Gambel oak, on limestone-derived or otherwise basic soils, often in rock crevices |
Elevation | 1500-3500 m (4900-11500 ft) | 1400-2900 (-3400) m (4600-9500 (-11200) ft) |
Distribution |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
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AZ; NM
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Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria pinetorum with reduced forms are found at high elevations; in disturbed, moist soils plants can become quite large, as in the Manzano Mountains. Densely cespitose plants with crowded racemes not exceeding the basal leaves are found at the crest (3200–3400 m) of the Sandia Mountains, New Mexico. These probably represent an undescribed taxon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 658. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | Lesquerella pinetorum |
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | (Wooton & Standley) O’Kane & Al-Shehbaz: Novon 12: 327. (2002) |
Web links |