Physaria acutifolia |
Physaria pendula |
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double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
Snake Range bladderpod |
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Habit | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | Perennials; caudex simple; densely pubescent, trichomes 5–7-rayed, rays bifurcate or trifurcate. |
Stems | several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
several from base, erect or outer usually decumbent toward base, (from below a terminal tuft of mostly erect leaves, unbranched), 1–2 dm. |
Basal leaves | (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
blade ovate to elliptic, 2–4 cm, (base gradually narrowed to petiole), margins entire, (surfaces densely pubescent with trichome layers). |
Cauline leaves | blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
(remote, proximal shortly petiolate); blade spatulate to oblanceolate, similar to basal, (base often cuneate), margins entire. |
Racemes | loose, (elongated in fruit). |
elongated. |
Flowers | sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
sepals linear-oblong, 5–7 mm, (lateral pair subsaccate); petals (erect), lingulate, 8–10 mm, (claw barely differentiated from blade). |
Fruiting pedicels | (divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
(recurved), 7–10 mm. |
Fruits | (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
(usually pendent, sessile), subglobose, slightly flattened (angustiseptate), 4–5 mm; valves densely pubescent, trichomes somewhat spreading; ovules 8–12 per ovary; style 4–5 mm. |
Seeds | (dark brown), flattened, (2–3 mm). |
plump, (mucilaginous when wetted). |
2n | = 10, 16, 24. |
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Physaria acutifolia |
Physaria pendula |
|
Phenology | Flowering May–Jun(-Jul). | Flowering May–Jun. |
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | Limestone gravel and cobbles, typically with junipers |
Elevation | 1500-3500 m (4900-11500 ft) | 1700-2100 m (5600-6900 ft) |
Distribution |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
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NV |
Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 657. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | Lesquerella pendula |
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 326. (2002) |
Web links |