Physaria acutifolia |
Physaria pallida |
|
---|---|---|
double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
white bladderpod |
|
Habit | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | Annuals (winter); with a fine taproot; sparsely pubescent, trichomes (minute), 3- or 4-rayed, rays furcate or, sometimes, trifurcate. |
Stems | several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
few to several from base, erect, (from basal leaf cluster, branched distally, flowering branches slender, subtended by bracts), 3–6 dm. |
Basal leaves | (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
blade oblanceolate or broadly obovate, to 10 cm, margins usually sinuate-dentate or entire, sometimes lobed. |
Cauline leaves | blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
(proximal shortly petiolate, distal sessile); blade oblanceolate to narrowly oblong, similar to basal, (distal with base slightly cuneate). |
Racemes | loose, (elongated in fruit). |
paniculate, (rachises and pedicels more densely pubescent than proximal leaves). |
Flowers | sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
sepals elliptic, 3–7 mm, (median pair slightly thickened apically, cucullate); petals (white), broadly ovate, to 12 mm, (narrowing gradually to short claw). |
Fruiting pedicels | (divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
(widely divaricate-ascending and straight, or slightly recurved), 10–15 mm, (slender, pubescent). |
Fruits | (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
(widely spreading to nearly pendent in age, shortly stipitate), globose or subglobose, not or slightly inflated, 3–10 mm; valves (not retaining seeds after dehiscence), glabrous; replum as wide as or wider than fruit; ovules 8–12 per ovary; style ca. 2 mm, (slender, fragile). |
Seeds | (dark brown), flattened, (2–3 mm). |
flattened. |
2n | = 10, 16, 24. |
= 12. |
Physaria acutifolia |
Physaria pallida |
|
Phenology | Flowering May–Jun(-Jul). | Flowering Apr–May. |
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | Grassy openings of small glade prairies, outcrops |
Elevation | 1500-3500 m (4900-11500 ft) | 90 m (300 ft) |
Distribution |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
|
TX |
Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria pallida is known from the Weches Formation in San Augustine County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 656. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | Vesicaria grandiflora var. pallida, Alyssum pallidum, Lesquerella pallida, Vesicaria pallida |
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | (Torrey & A. Gray) O’Kane & Al-Shehbaz: Novon 12: 326. (2002) |
Web links |