Physaria acutifolia |
Physaria occidentalis |
|||||
---|---|---|---|---|---|---|
double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
western bladder-pod |
|||||
Habit | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | Perennials; caudex simple or branched, (thickened, sometimes subterranean); densely pubescent, trichomes (sessile or short-stalked), 5–7-rayed, raysdistinct, bifurcate, (moderately to prominently tuberculate). | ||||
Stems | several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
few to several from base, prostrate to decumbent or erect, (usually unbranched), 0.3–1.5(–3) dm. |
||||
Basal leaves | (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
(petiole slender); blade suborbicular to obovate or elliptic, 1–8 cm, (base narrowing gradually or abruptly to petiole), margins sinuate-dentate or entire. |
||||
Cauline leaves | blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
(proximal shortly petiolate, distal sessile); blade oblanceolate, 0.5–1.5(–2.5) cm, margins entire or remotely dentate. |
||||
Racemes | loose, (elongated in fruit). |
dense or loose. |
||||
Flowers | sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
sepals elliptic or oblong-elliptic, 4.5–7 mm, (median pair cucullate); petals spatulate, 7–9(–14) mm, (claw undifferentiated from blade). |
||||
Fruiting pedicels | (divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
(sigmoid or curved), 5–10(–15) mm. |
||||
Fruits | (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
(erect), ellipsoid to obovoid, compressed at apex and sometimes margins (strongly latiseptate), (5–)6–9 mm, (apex acute, often beaked); valves densely pubescent, trichomes appressed or spreading, sometimes sparsely pubescent inside, trichomes 4- or 5-rayed; ovules 4–12 per ovary; style (2–)3–6.5 mm, (often sparsely pubescent). |
||||
Seeds | (dark brown), flattened, (2–3 mm). |
inner surface flattened, outer convex. |
||||
2n | = 10, 16, 24. |
|||||
Physaria acutifolia |
Physaria occidentalis |
|||||
Phenology | Flowering May–Jun(-Jul). | |||||
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | |||||
Elevation | 1500-3500 m (4900-11500 ft) | |||||
Distribution |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
|
CA; ID; NV; OR; UT; WA
|
||||
Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||
Key |
|
|||||
Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 654. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | Vesicaria occidentalis, Lesquerella occidentalis | ||||
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | (S. Watson) O’Kane & Al-Shehbaz: Novon 12: 326. (2002) | ||||
Web links |