Physaria acutifolia |
Physaria obdeltata |
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double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
Middle Butte bladderpod |
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Habit | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | Perennials; caudex simple; densely pubescent, trichomes (appressed in layers), 5–7-rayed, rays usually bifurcate, sometimes furcate, (thickened toward center). |
Stems | several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
simple from base, prostrate to decumbent, (unbranched, from within and below leaf clusters, slender), 0.2–0.8 dm. |
Basal leaves | (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
(tufted, erect or ascending, silvery); blade linear to oblanceolate or (outer) oblanceolate to obovate or rhombic, 1.5–3.3 cm, (base sometimes subhastate), margins entire or dentate, (often involute). |
Cauline leaves | blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
blade nearly linear, to 1.5 cm, margins entire. |
Racemes | loose, (elongated in fruit). |
dense, (subcorymbose). |
Flowers | sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
sepals (yellow-green), lanceolate, (2.5–)3.5–4.5 mm; petals spatulate to oblanceolate, 4–6.5 mm. |
Fruiting pedicels | (divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
(recurved), 5–8 mm. |
Fruits | (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
(usually pendent), obdeltate, compressed (angustiseptate), 2–4 mm, (wider than long, base tapered to acute angle, apex truncate with inflated shoulders); valves densely pubescent; (septum obsolete or with a narrow fringe inside of replum, funicles attached close to replum apex); ovules 4 per ovary; style 2–4 mm, (slender). |
Seeds | (dark brown), flattened, (2–3 mm). |
plump, (mucilaginous when wetted). |
2n | = 10, 16, 24. |
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Physaria acutifolia |
Physaria obdeltata |
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Phenology | Flowering May–Jun(-Jul). | Flowering Jun(-Jul). |
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | Clayey, silty, or gravelly soils, overlaying basalt lava flows, silty playas, sagebrush, barren areas |
Elevation | 1500-3500 m (4900-11500 ft) | 1300-1700 m (4300-5600 ft) |
Distribution |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
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ID |
Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria obdeltata is known from the eastern Snake River Plain. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 654. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | Lesquerella obdeltata |
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 326. (2002) |
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