Physaria acutifolia |
Physaria multiceps |
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double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
manyhead bladderpod |
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Habit | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | Perennials; caudex (buried), branched, (not thickened); densely pubescent, trichomes (sessile or short-stalked), 5–8-rayed, rays furcate or bifurcate, (umbonate, rough to finely tuberculate throughout). |
Stems | several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
several from base, prostrate, (slender, sparsely pubescent), 0.5–2 dm. |
Basal leaves | (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
blade obovate to narrowly elliptic, 1.5–6 cm, margins usually entire, rarely shallowly dentate, (surfaces densely pubescent, often silvery). |
Cauline leaves | blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
blade oblanceolate to spatulate, 0.5–1 cm, margins entire, (surfaces often sparsely pubescent). |
Racemes | loose, (elongated in fruit). |
(narrow), loose, (elongated in fruit). |
Flowers | sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
sepals (greenish brown, sometimes magenta), linear or elliptic, 4.3–6(–7.5) mm, (median pair thickened apically, cucullate); petals (frequently pink or magenta in distal 1/3–1/2), spatulate to oblanceolate, 6–10(–12) mm, (claw undifferentiated from blade). |
Fruiting pedicels | (divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
(ascending to somewhat spreading, straight to slightly curved), 4–8(–12) mm. |
Fruits | (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
broadly ovoid to suborbicular, inflated, (terete or, often, slightly angustiseptate), 3–6mm; valves sparsely pubescent; ovules usually 4, rarely 6–8 per ovary; style 3–6.5 mm. |
Seeds | (dark brown), flattened, (2–3 mm). |
plump. |
2n | = 10, 16, 24. |
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Physaria acutifolia |
Physaria multiceps |
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Phenology | Flowering May–Jun(-Jul). | Flowering May–Jul. |
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | Douglas-fir or spruce woodlands, limestone ridges, damp open slopes, soil pockets among rocks, crevices of rocks, decomposed calcareous rocks |
Elevation | 1500-3500 m (4900-11500 ft) | 2400-2900 m (7900-9500 ft) |
Distribution |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
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ID; UT; WY |
Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 651. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | Lesquerella multiceps |
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | (Maguire) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) |
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