Physaria acutifolia |
Physaria macrocarpa |
|
---|---|---|
double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
largefruit bladderpod |
|
Habit | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | Perennials; caudex branched; densely pubescent, trichomes (sessile or short-stalked), 4–6-rayed, rays distinct, usually furcate, rarely bifurcate, (finely tuberculate throughout). |
Stems | several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
few or several from base, prostrate to decumbent, (unbranched or branched), 0.5–1.5 dm. |
Basal leaves | (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
blades orbicular to broadly obovate, 1.5–3 cm, margins usually entire, rarely remotely dentate. |
Cauline leaves | blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
(sessile or shortly petiolate); blade elliptic to oblanceolate, 1–1.5(–2.5) cm, margins entire, (apex obtuse). |
Racemes | loose, (elongated in fruit). |
dense, (elongated in fruit). |
Flowers | sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
sepals ovate or oblong-elliptic, 5–5.5 mm, (lateral pair not saccate); petals cuneate or broadly obovate, ca. 7 mm, (sometimes slightly narrowed to a broad claw, apex sometimes retuse). |
Fruiting pedicels | (divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
(sharply recurved), 5–10 mm, (stout). |
Fruits | (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
subglobose to broadly obovoid, strongly inflated (often slightly angustiseptate), 5–7 mm, (papery); valves sparsely pubescent; (septum fenestrate, perforate, or obsolete); ovules 4–8 per ovary; style 2–3 mm. |
Seeds | (dark brown), flattened, (2–3 mm). |
somewhat flattened. |
2n | = 10, 16, 24. |
|
Physaria acutifolia |
Physaria macrocarpa |
|
Phenology | Flowering May–Jun(-Jul). | Flowering May–Jun. |
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | Gypsum-clay hills and benches, naked clay flats and barren hills |
Elevation | 1500-3500 m (4900-11500 ft) | 2000-2400 m (6600-7900 ft) |
Distribution |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
|
WY |
Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria macrocarpa is found in the Great Divide and Green River basins. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 650. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | Lesquerella macrocarpa |
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | (A. Nelson) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) |
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