Physaria acutifolia |
Physaria lepidota |
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double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
Kane County twinpod |
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Habit | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | Perennials; caudex simple, (with deep roots, thickened); densely (silvery) pubescent throughout (densely covering leaves with several appressed layers), less dense on stems, trichomes (stellate-scalelike), rays fused (webbed) in proximal 1/2 or to tips, (umbonate, nearly smooth to moderately tuberculate). | ||||
Stems | several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
simple from base, erect or outer ones slightly decumbent toward base, (from below or in basal leaves, unbranched), (0.5–)0.8–1.6(–2) dm. |
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Basal leaves | (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
(erect, petiole long, slender); blade spatulate to broadly oblanceolate, (3–)5–7(–12) cm, (base gradually tapering to petiole), margins entire, (apex rounded or obtuse). |
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Cauline leaves | blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
blade oblanceolate, similar to basal, (base cuneate), margins entire. |
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Racemes | loose, (elongated in fruit). |
dense. |
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Flowers | sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
sepals (erect), linear to linear-oblong, somewhat boat-shaped, 7–10 mm; petals (erect at anthesis), lingulate, 11–15 mm, (claw undifferentiated from blade). |
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Fruiting pedicels | (divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
(divaricate-ascending, straight or slightly curved), 10–15 mm. |
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Fruits | (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
(purplish in age), strongly didymous, semiorbicular, highly inflated, 10–18 × 14–19 mm, (papery), basal sinus usually shallow, rarely absent, apical sinus deep, narrowly V-shaped; valves retaining seeds after dehiscence, sides flat, back rounded, margins keeled, base and apex obtuse; replum narrowly oblong to linear, as wide as or wider than fruit, base slightly narrowed, apex obtusely rounded; ovules 4 per ovary; style 3–5 mm, (slender). |
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Seeds | (dark brown), flattened, (2–3 mm). |
slightly flattened. |
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2n | = 10, 16, 24. |
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Physaria acutifolia |
Physaria lepidota |
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Phenology | Flowering May–Jun(-Jul). | |||||
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | |||||
Elevation | 1500-3500 m (4900-11500 ft) | |||||
Distribution |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
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UT |
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Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 648. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | |||||
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | Rollins: Brittonia 33: 335, figs. 1, 2. (1981) | ||||
Web links |