Physaria acutifolia |
Physaria klausii |
|
---|---|---|
double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
Rogers Pass or klaus' or divide bladderpod |
|
Habit | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | Perennials; caudex simple; densely pubescent, trichomes (loosely spreading), 3–5-rayed, rays distinct, furcate (with exceptionally long branches). |
Stems | several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
simple from base, erect to decumbent, (slender), 0.6–1.5 dm. |
Basal leaves | (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
blades obovate to deltate, 1.5–3(–4) cm, margins entire or outer one with 1 or 2 broad teeth. |
Cauline leaves | blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
blade oblanceolate to spatulate, 0.6–1.5 cm, margins entire. |
Racemes | loose, (elongated in fruit). |
loose. |
Flowers | sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
sepals (green-yellow, often tinged with purple), elliptic, 3–4.6 mm; petals oblanceolate, 6–8 mm (claw expanded). |
Fruiting pedicels | (divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
(sigmoid), 5–9 mm. |
Fruits | (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
(depressed), broadly obovate, compressed (angustiseptate), 2–4 mm, (apex slightly bilobed to nearly truncate); valves densely pubescent, trichomes strongly ascending, spreading, long, (appearing fuzzy), pubescent inside; ovules 4 per ovary; style 3–4 mm, (pubescent or glabrous). |
Seeds | (dark brown), flattened, (2–3 mm). |
flattened. |
2n | = 10, 16, 24. |
|
Physaria acutifolia |
Physaria klausii |
|
Phenology | Flowering May–Jun(-Jul). | Flowering Jul. |
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | Open gravel slides, solifluction cross-stripes of shale rubble, barren shale-derived soil |
Elevation | 1500-3500 m (4900-11500 ft) | 1200-1900 m (3900-6200 ft) |
Distribution |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
|
MT |
Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 647. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | Lesquerella klausii |
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | (Rollins) O’Kane & Al-Shehbaz: Novon 12: 325. (2002) |
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