The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod

Hitchcock's bladderpod

Habit Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). Perennials; (forming loose mats); caudex (buried), branched; densely pubescent, trichomes (short-stalked), 4–6-rayed, rays distinct, bifurcate, (rough-tuberculate).
Stems

several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm.

few to several from base, prostrate to erect or spreading, 0.05–0.5(–1.2) dm.

Basal leaves

(petiole slender, often narrowly winged);

blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro).

(petiole and blade differentiated or not);

blade spatulate to elliptic or linear or narrowly oblanceolate, 0.5–1.5(–2.5) cm, margins entire.

Cauline leaves

blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse).

similar to basal, smaller.

Racemes

loose, (elongated in fruit).

dense.

Flowers

sepals linear-oblong, 4–7.5 mm;

petals spatulate, 6–11 mm.

sepals narrowly lanceolate to lanceolate, 2.8–6 mm;

petals (pale to deep yellow), narrowly lanceolate to oblanceolate, 5–9 mm, (claw undifferentiated from blade).

Fruiting pedicels

(divaricate, slightly sigmoid or nearly straight), 6–12 mm.

(ascending, straight or slightly curved), 2–6 mm.

Fruits

(erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so);

valves retaining seeds after dehiscence, pubescent, trichomes appressed;

replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse;

ovules (2 or) 4 per ovary;

style 4–6(–9) mm.

(sessile or substipitate), globose or subglobose to obovoid, not or slightly inflated, 3–6 mm, (firm, apex acute);

valves (reddish in age, not retaining seeds after dehiscence), glabrous throughout;

replum as wide as or wider than fruit;

ovules 4–8 per ovary;

style 1.7–6 mm.

Seeds

(dark brown), flattened, (2–3 mm).

flattened.

2n

= 10, 16, 24.

Physaria acutifolia

Physaria hitchcockii

Phenology Flowering May–Jun(-Jul).
Habitat Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities
Elevation 1500-3500 m (4900-11500 ft)
Distribution
from FNA
AZ; CO; ID; MT; NM; NV; SD; UT; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
NV; UT
[BONAP county map]
Discussion

Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 3 (3 in the flora).

The taxonomic treatment of Physaria hitchcockii has varied widely over the years. Molecular study (pers. obs.) has shown no direct relationship to P. tumulosa; morphologically, though, P. navajoensis and P. tumulosa appear closely related. Infraspecific taxonomy is based on the presence of a discernable petiole and whether or not the caudex is elastically elongated. The subspecies recognized here are usually geographically coherent, except that collections from the Table Cliff Plateau are more similar to subsp. hitchcockii, disjunct in Nevada, than they are to the very nearly sympatric subsp. rubicundula.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Caudices elongated and elastic; basal leaves: petiole not differentiated from blade, blades linear-oblanceolate; Aquarius, Markagunt, and Paunsaugunt plateaus, Utah (limited to the pink member of the limestone Wasatch (Claron) Formation).
subsp. rubicundula
1. Caudices elongated or not, elastic or not; basal leaves: petiole differentiated (sometimes weakly) from blade, blades oblanceolate to obovate; Nevada, Utah
→ 2
2. Plants forming tufts; caudices not elongated, not elastic; fruits 2.6-3.8 mm wide; Table Cliff Plateau, Utah (limited to the white member of the limestone Wasatch (Claron) Formation) and limestones of the Sheep Range and Spring Mountains, Nevada.
subsp. hitchcockii
2. Plants forming soft mats; caudices elongated, elastic (creeping); fruits 1.7-3 mm wide; Grant, Quinn Canyon, and Schell Creek ranges, Nevada.
subsp. confluens
Source FNA vol. 7, p. 624. FNA vol. 7, p. 643.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Subordinate taxa
P. hitchcockii subsp. confluens, P. hitchcockii subsp. hitchcockii, P. hitchcockii subsp. rubicundula
Synonyms P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa Lesquerella hitchcockii
Name authority Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) (Munz) O’Kane & Al-Shehbaz: Novon 12: 324. (2002)
Web links