Physaria acutifolia |
Physaria geyeri |
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double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
Geyer's bladderpod, Geyer's twin-pod |
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Habit | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | Perennials; caudex usually simple, (cespitose); (silvery) pubescent throughout, trichomes (sessile), 6–8-rayed, rays mostly furcate, (tuberculate to nearly smooth). | ||||
Stems | several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
several from base, decumbent, (arising laterally, unbranched), 1–3 dm. |
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Basal leaves | (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
(numerous); (petiole slender, rarely with a few broad teeth); blade obovate, 3–7 cm, margins entire. |
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Cauline leaves | blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
blade oblanceolate, 1.5–3 cm, margins entire. |
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Racemes | loose, (elongated in fruit). |
loose. |
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Flowers | sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
sepals oblong, 5–7 mm; petals (yellow to purplish), spatulate, 8–12 mm. |
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Fruiting pedicels | (divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
(ascending or spreading, slightly curved or sigmoid), 1–2 cm. |
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Fruits | (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
obcordate, angustiseptate, somewhat inflated, (not bladdery), 5–7 × 6–9 mm, (papery, basal sinus absent, apical sinus broad and open); valves (retaining seeds after dehiscence), loosely pubescent, trichomes spreading; replum ovate, 5–7 mm, as wide as or wider than fruit, apex acute or obtuse; ovules 4–6 per ovary; style 5–7 mm. |
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Seeds | (dark brown), flattened, (2–3 mm). |
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2n | = 10, 16, 24. |
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Physaria acutifolia |
Physaria geyeri |
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Phenology | Flowering May–Jun(-Jul). | |||||
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | |||||
Elevation | 1500-3500 m (4900-11500 ft) | |||||
Distribution |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
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ID; MT; OR; WA
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Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 638. | ||||
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | Vesicaria geyeri, Coulterina geyeri, Lesquerella geyeri | ||||
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | (Hooker) A. Gray: Gen. Amer. Bor. 1: 162. (1848) | ||||
Web links |