Physaria acutifolia |
Physaria fremontii |
|
---|---|---|
double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
Fremont's bladderpod |
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Habit | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | Perennials; caudex simple; densely pubescent, trichomes (subsessile), 5–7-rayed, rays distinct or slightly fused at base, usually furcate, (roughly tuberculate). |
Stems | several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
few to several from base, prostrate, (arising proximal to a terminal cluster of erect leaves, usually unbranched, slender), 0.5–1(–1.5) dm. |
Basal leaves | (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
(petiole slender); blade elliptic to rhombic, 1.5–4(–5) cm, (base gradually tapering to petiole), margins usually entire, rarely dentate. |
Cauline leaves | blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
blade narrowly obovate to oblanceolate, 5–15 mm, margins entire. |
Racemes | loose, (elongated in fruit). |
(relatively short), rather loose. |
Flowers | sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
sepals ± elliptic, 4–6 mm; petals ovate, 6–8 mm, (claw slightly expanded at base). |
Fruiting pedicels | (divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
(usually secund, recurved), 5–8 mm. |
Fruits | (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
(pendent), globose, subglobose, or slightly obcompressed, slightly or not inflated, 3–6 mm, (rigid, apex usually beaked); valves pubescent throughout; ovules 8–12 per ovary; style 1–2 mm, (pubescent). |
Seeds | (dark brown), flattened, (2–3 mm). |
slightly flattened, (suborbicular). |
2n | = 10, 16, 24. |
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Physaria acutifolia |
Physaria fremontii |
|
Phenology | Flowering May–Jun(-Jul). | Flowering May–Jun. |
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | Calcareous gravel, loose whitish rubble, limestone pavement, rocky calcareous ridges |
Elevation | 1500-3500 m (4900-11500 ft) | 2100-2800 m (6900-9200 ft) |
Distribution |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
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WY |
Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria fremontii is known from the area of the Wind River Mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 638. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | Lesquerella fremontii |
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | (Rollins & E. A. Shaw) O’Kane & Al-Shehbaz: Novon 12: 323. (2002) |
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