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double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod

Dorn's twinpod, tunp range twinpod

Habit Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). Perennials; (compact); caudex simple, (stout); densely (silvery) pubescent throughout (except style), trichomes several-rayed, rays furcate, fused at base, (umbonate, tuberculate throughout).
Stems

several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm.

simple from base, erect, (arising from a condensed rosette), to 1 dm.

Basal leaves

(petiole slender, often narrowly winged);

blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro).

(ascending or erect);

blade elliptic to oblanceolate to obovate, (usually curled from middle to apex), (1.5–)5–7 cm (width 12–20 mm), margins entire.

Cauline leaves

blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse).

(1–5);

blade oblanceolate, 1–2.5 cm, margins entire.

Racemes

loose, (elongated in fruit).

compact (or elongated in fruit, to 1 dm, barely exceeding leaves).

Flowers

sepals linear-oblong, 4–7.5 mm;

petals spatulate, 6–11 mm.

sepals (erect), oblong to linear or spatulate, 5.5–7 mm;

petals spatulate, 10–14 mm, (claw undifferentiated from blade).

Fruiting pedicels

(divaricate, slightly sigmoid or nearly straight), 6–12 mm.

(divaricate-ascending, slightly curved), 7–18 mm.

Fruits

(erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so);

valves retaining seeds after dehiscence, pubescent, trichomes appressed;

replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse;

ovules (2 or) 4 per ovary;

style 4–6(–9) mm.

didymous, irregular, highly inflated, 8–11(–18) × 10–15 mm, (papery, basal sinus shallower than the deep apical sinus; valves retaining seeds after dehiscence);

replum obovate, not constricted, 1–1.8 mm, apex obtuse, as wide as or wider than fruit;

ovules (4–)8(–12) per ovary;

style 4–6 mm, (glabrous).

Seeds

(dark brown), flattened, (2–3 mm).

flattened, (oblong to elliptic, thin-margined or not).

2n

= 10, 16, 24.

Physaria acutifolia

Physaria dornii

Phenology Flowering May–Jun(-Jul). Flowering May–Jun.
Habitat Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities Calcareous shale, slopes, ridges
Elevation 1500-3500 m (4900-11500 ft) 1900-2200 m (6200-7200 ft)
Distribution
from FNA
AZ; CO; ID; MT; NM; NV; SD; UT; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
WY
[BONAP county map]
Discussion

Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 624. FNA vol. 7, p. 634.
Parent taxa Brassicaceae > tribe Physarieae > Physaria Brassicaceae > tribe Physarieae > Physaria
Sibling taxa
P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. dornii, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
P. acutifolia, P. alpestris, P. alpina, P. angustifolia, P. arctica, P. arenosa, P. argyraea, P. arizonica, P. aurea, P. bellii, P. brassicoides, P. calcicola, P. calderi, P. carinata, P. chambersii, P. cinerea, P. condensata, P. congesta, P. cordiformis, P. curvipes, P. densiflora, P. didymocarpa, P. douglasii, P. eburniflora, P. engelmannii, P. eriocarpa, P. fendleri, P. filiformis, P. floribunda, P. fremontii, P. garrettii, P. geyeri, P. globosa, P. gooddingii, P. gordonii, P. gracilis, P. grahamii, P. hemiphysaria, P. hitchcockii, P. humilis, P. integrifolia, P. intermedia, P. kingii, P. klausii, P. lata, P. lepidota, P. lesicii, P. lindheimeri, P. ludoviciana, P. macrocarpa, P. mcvaughiana, P. montana, P. multiceps, P. navajoensis, P. nelsonii, P. newberryi, P. obcordata, P. obdeltata, P. occidentalis, P. oregona, P. ovalifolia, P. pachyphylla, P. pallida, P. parviflora, P. parvula, P. pendula, P. pinetorum, P. prostrata, P. pruinosa, P. pulvinata, P. purpurea, P. pycnantha, P. rectipes, P. recurvata, P. reediana, P. rollinsii, P. saximontana, P. scrotiformis, P. sessilis, P. spatulata, P. subumbellata, P. tenella, P. thamnophila, P. tumulosa, P. valida, P. vicina, P. vitulifera
Synonyms P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa
Name authority Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) Lichvar: Brittonia 35: 150, figs. 1–3. (1983)
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