Physaria acutifolia |
Physaria condensata |
|
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double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
tuft twinpod |
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Habit | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | Perennials; caudex usually simple, rarely branched, (enlarged with persistent leaf bases, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays typically furcate, (fused at base, arms slender, tuberculate throughout). |
Stems | several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
several from base, decumbent to ascending, (arising laterally beneath a dense rosette), less than 0.1 dm. |
Basal leaves | (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
(petiole slender); blade (horizontal on the ground), obovate, 0.5–1.5 cm (width 4–8 mm, base tapering abruptly to petiole), margins entire, (apex usually acute, surfaces silvery from a dense incrustation of appressed, stellate trichomes). |
Cauline leaves | blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
blade oblanceolate, 0.5–1 cm (width 2–3 mm), margins entire, (surfaces densely stellate pubescent). |
Racemes | loose, (elongated in fruit). |
congested, (subumbellate, often almost sessile, barely exceeding basal leaves). |
Flowers | sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
sepals (yellowish green), narrowly lanceolate, 4–5 mm; petals (erect), oblanceolate, 6–7 mm, (claw weakly differentiated from blade). |
Fruiting pedicels | (divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
(divaricate, straight), 5–10 mm. |
Fruits | (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
didymous, ovate, inflated, 4.8–6 × 6–10 mm, (papery, basal and apical sinuses deep); valves (retaining seeds after dehiscence), pubescent, trichomes loosely spreading; replum obovate, 3–4 mm, as wide as or wider than fruit; ovules 8 per ovary (2–4 abortive); style 4–6 mm. |
Seeds | (dark brown), flattened, (2–3 mm). |
flattened. |
2n | = 10, 16, 24. |
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Physaria acutifolia |
Physaria condensata |
|
Phenology | Flowering May–Jun(-Jul). | Flowering May–Jun. |
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | Calcareous knolls and ridges, clay banks, limey slopes, shaley hills, clay patches |
Elevation | 1500-3500 m (4900-11500 ft) | 1800-2400 m (5900-7900 ft) |
Distribution |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
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WY |
Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 631. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | |
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | Rollins: Rhodora 41: 407, plate 556, figs. 1, 9, 10. (1939) |
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