Physaria acutifolia |
Physaria calderi |
|
---|---|---|
double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
Calder's bladderpod |
|
Habit | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | Perennials; caudex simple or branched; densely pubescent throughout, trichomes (sessile or subsessile), rays distinct or slightly fused at base, furcate or bifurcate, (strongly umbonate, tuberculate, tubercles often relatively larger, fewer over center). |
Stems | several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
simple or few to several from base, usually erect to spreading, sometimes prostrate, 0.5–2 dm. |
Basal leaves | (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
blade oblanceolate, 2–3 cm, margins entire. |
Cauline leaves | blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
(sessile or proximal shortly petiolate); blade narrowly oblanceolate, 0.5–1.5 cm, margins entire. |
Racemes | loose, (elongated in fruit). |
loose. |
Flowers | sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
sepals ovate to elliptic, (3–)4–5(–6) mm, (median pair often thickened apically, cucullate); petals obovate, (6–)7–10 mm (nearly as wide, abruptly narrowed to claw, ca. 1 mm wide). |
Fruiting pedicels | (divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
(erect to divaricate or ascending, sometimes curved), (5–)10–20(–40) mm, (stout). |
Fruits | (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
subglobose to ellipsoid, compressed (usually angustiseptate), to 8 mm; (valves not retaining seeds after dehiscence); replum as wide as or wider than fruit; ovules 10–14 per ovary; style 1–2 mm. |
Seeds | (dark brown), flattened, (2–3 mm). |
plump. |
2n | = 10, 16, 24. |
= 20. |
Physaria acutifolia |
Physaria calderi |
|
Phenology | Flowering May–Jun(-Jul). | Flowering Jun–Aug. |
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | Dry rocky summits, limestone flats and slopes, alpine knolls |
Elevation | 1500-3500 m (4900-11500 ft) | 600-1500 m (2000-4900 ft) |
Distribution |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
|
AK; NT; YT |
Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physaria calderi is known from the Ogilvie and Richardson mountains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 629. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | Lesquerella calderi, Lesquerella arctica subsp. calderi |
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | (G. A. Mulligan & A. E. Porsild) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) |
Web links |