Physaria acutifolia |
Physaria aurea |
|
---|---|---|
double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
golden bladderpod |
|
Habit | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | Biennials or perennials; (short-lived); caudex branched; densely pubescent, trichomes (sessile or short-stalked, simple or not), 5–9-rayed, rays furcate, (fine, smooth or finely tuberculate). |
Stems | several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
several from base, erect or outer ones decumbent or procumbent, (sometimes much-branched distally), to 6 dm. |
Basal leaves | (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
blade obovate or rhombic, to ca. 2.5 cm, margins usually shallowly dentate, sometimes lyrate-pinnatifid. |
Cauline leaves | blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
(proximal shortly petiolate, distal sessile); blade obovate to rhombic or oblanceolate, 2–4(–6) cm, margins entire or shallowly and remotely dentate. |
Racemes | loose, (elongated in fruit). |
usually dense, (several-flowered). |
Flowers | sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
sepals ovate or oblong (tapering at base), 3.6–4.8(–5.3) mm, (lateral pair subsaccate, median pair thickened apically, cucullate); petals obovate to spatulate, 4.5–7.5 mm, (blade narrowed to broad claw, margins sinuate). |
Fruiting pedicels | (divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
(strongly recurved), to 20 mm. |
Fruits | (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
(± pendent), ovoid, obcompressed, or globose, compressed, 4–6(–8) mm; valves (not retaining seeds after dehiscence), sparsely pubescent or glabrous, sparsely pubescent inside; replum as wide as or wider than fruit; ovules usually 4, rarely 6, per ovary; style 2.5–3.6 mm. |
Seeds | (dark brown), flattened, (2–3 mm). |
flattened. |
2n | = 10, 16, 24. |
= 14. |
Physaria acutifolia |
Physaria aurea |
|
Phenology | Flowering May–Jun(-Jul). | Flowering Jun–Aug. |
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | Open sites and bare areas in rocky limestone soil in mountains, roadbanks, open woods |
Elevation | 1500-3500 m (4900-11500 ft) | 2000-2800 m (6600-9200 ft) |
Distribution |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
|
NM |
Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Physaria aurea (known from the Jicarilla and Sacramento mountains) is similar to 35. P. gooddingii, which is found farther west in the mountains of Catron and Sierra counties, New Mexico, and Greenlee County, Arizona. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 628. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | Lesquerella aurea |
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | (Wooton) O’Kane & Al-Shehbaz: Novon 12: 322. (2002) |
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