Physaria acutifolia |
Physaria angustifolia |
|
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double bladderpod, pointleaf twinpod, Rydberg's twinpod, sharpleaf twinpod |
threadleaf bladderpod |
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Habit | Perennials; caudex branched, (sometimes forming a thick crown, cespitose); (silvery) pubescent throughout, trichomes several-rayed, rays furcate, (moderately tuberculate, rays weakly so). | Annuals; with a fine taproot; ± densely pubescent, trichomes several-rayed, rays distinct or fused at base, bifurcate, (prominently tuberculate throughout). |
Stems | several from base, usually somewhat decumbent, (unbranched), (0.4–)0.5–2 dm. |
simple or few to several from base, erect, (sometimes branched), to 4 dm. |
Basal leaves | (petiole slender, often narrowly winged); blade obovate to orbicular or rhombic-orbicular, 2–9 cm, (base abruptly narrowed to petiole), margins usually entire, rarely with few scattered teeth, (apex rounded or obtuse, sometimes with apical mucro). |
blade elliptic to rhombic, 3–8 cm, (base narrowing gradually to petiole), margins entire, repand, coarsely toothed, or pinnatifid. |
Cauline leaves | blade spatulate to oblanceolate, 1–3 cm, margins entire, (apex usually obtuse). |
(proximal often shortly petiolate, distal sessile); blade linear or narrowly obovate, 1.5–6(–10) cm, margins entire, repand, or shallowly toothed. |
Racemes | loose, (elongated in fruit). |
usually loose. |
Flowers | sepals linear-oblong, 4–7.5 mm; petals spatulate, 6–11 mm. |
sepals elliptic or ovate, 4–6 mm, (lateral pair usually subsaccate); petals obovate to obdeltate, 6–10 mm, (apex often emarginate). |
Fruiting pedicels | (divaricate, slightly sigmoid or nearly straight), 6–12 mm. |
(usually divaricate, sometimes horizontal, straight or slightly curved), 8–20 mm. |
Fruits | (erect), didymous, suborbicular, inflated, (4–)6–15 × (4–)8–20 mm, (papery, basal and apical sinuses similar, basal rarely shallower, apical deep, narrow and closed or nearly so); valves retaining seeds after dehiscence, pubescent, trichomes appressed; replum oblong, constricted, 2–3.5 mm, narrower than fruit, apex obtuse; ovules (2 or) 4 per ovary; style 4–6(–9) mm. |
not didymous, ± globose, slightly inflated, 4–6 mm; valves (not retaining seeds after dehiscence), glabrous throughout; replum as wide as or wider than fruit; ovules 4 per ovary; style 2–3.5 mm; (stigma expanded). |
Seeds | (dark brown), flattened, (2–3 mm). |
flattened, (margined). |
2n | = 10, 16, 24. |
= 10. |
Physaria acutifolia |
Physaria angustifolia |
|
Phenology | Flowering May–Jun(-Jul). | Flowering Apr(-May). |
Habitat | Hillsides, roadcuts, sagebrush, pinyon-juniper, Gambel oak, ponderosa pine communities | Shallow limestone-derived soils, sometimes spreading to disturbed sites |
Elevation | 1500-3500 m (4900-11500 ft) | 90-300 m (300-1000 ft) |
Distribution |
AZ; CO; ID; MT; NM; NV; SD; UT; WY
|
OK; TX |
Discussion | Physaria acutifolia tends to be somewhat dwarfed, with a branched caudex and especially long styles (var. stylosa), where it grows at high elevations, especially at the western end of the Uinta Mountains in Utah. Intermediates form an uninterrupted cline and no infraspecific taxa are here recognized. In R. C. Rollins (1939), the discussion of P. acutifolia actually pertains to P. rollinsii. The discussion of P. australis pertains to what is now known as P. acutifolia. The plants are usually found in open soil patches, rarely into the subalpine or alpine tundra. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 7, p. 624. | FNA vol. 7, p. 625. |
Parent taxa | Brassicaceae > tribe Physarieae > Physaria | Brassicaceae > tribe Physarieae > Physaria |
Sibling taxa | ||
Synonyms | P. acutifolia var. stylosa, P. australis, P. didymocarpa var. australis, P. stylosa | Vesicaria angustifolia, Lesquerella angustifolia, Lesquerella longifolia |
Name authority | Rydberg: Bull. Torrey Bot. Club 28: 279. (1901) | (Nuttall ex Torrey & A. Gray) O’Kane & Al-Shehbaz: Novon 12: 321. (2002) |
Web links |