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hairy ground-cherry, husk tomato

ivy leafed tomatillo, ivyleaf groundcherry

Habit Herbs annual, taprooted, ± glabrous to villous, hairs simple, jointed, glandular and eglandular, of varying lengths, all shorter than 0.5 mm, plants from southwestern United States all glandular, green in appearance when dry. Herbs perennial, rhizomatous, rhizome stout, densely pubescent, hairs simple, sometimes jointed, glandular, 0.5–1 mm, sometimes also with sessile glands.
Stems

erect, branching at most nodes, branches spreading, 0.5–8 dm.

erect to decumbent, usually branching from base and at most nodes, branches spreading, 0.5–3 dm.

Leaves

petiolate;

petiole 1/5 to as long as blade;

blade broadly ovate to orbiculate, (1.6–)2.5–8(–9.5) × (1–)2–7 cm, base rounded to slightly cordate, margins entire or coarsely dentate, teeth fewer than 8 per side.

petiolate;

petiole 1/2 to ± as long as blade;

blade broadly ovate to orbiculate, 1.5–3.5 × 1–3 cm, base cordate to rounded, margins ± entire or coarsely dentate, teeth sharp to blunt.

Pedicels

slender, 3.5–9 mm, 5–15 mm in fruit.

4–8(–13) mm, 5–15 mm in fruit.

Flowers

calyx 3–6(–7) mm, lobes 1–3.5 mm;

corolla yellow with 5 large, dark purple-brown-black spots, campanulate-rotate, 6–11 mm;

anthers blue, rarely yellow or blue-tinged, not twisted after dehiscence, 1–2 mm.

calyx 5–7(–10) mm, lobes 1.5–3.5(–5) mm;

corolla yellow with 5 dark brown spots, campanulate-rotate, 7–12 mm;

anthers yellow, not twisted after dehiscence, 2–4 mm.

Fruiting calyces

loosely enclosing berry, sharply 5-angled, 20–35 × 15–25(–30) mm, always noticeably longer than wide.

loosely enclosing berry, 10-ribbed, 20–30 × 15–25(–30) mm.

2n

= 24.

= 24.

Physalis pubescens

Physalis hederifolia

Phenology Flowering year-round in areas without frost, mostly May–Oct. Flowering Apr–Aug.
Habitat Low woods, edges of swamps, stream banks, floodplains, hammocks, disturbed habitats. Dry open gravelly sites, rocky ledges, open plains.
Elevation 0–900 m. [0–3000 ft.] 200–2600 m. [700–8500 ft.]
Distribution
from FNA
AL; AR; AZ; CA; DC; FL; GA; IA; IL; IN; KS; KY; LA; MD; MI; MO; MS; NC; NJ; NM; OH; OK; PA; SC; TN; TX; UT; VA; WI; WV; Mexico; Central America; South America; West Indies [Introduced in Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; KS; MT; NE; NM; NV; OK; SD; TX; UT; WY; Mexico (Baja California, Chihuahua, Coahuila, Durango, Nuevo León, San Luis Potosí, Sonora, Tamaulipas, Zacatecas)
[WildflowerSearch map]
[BONAP county map]
Discussion

Fresh plants of Physalis pubescens reportedly have a strong fetid odor (M. Martínez 1998). This widespread species exhibits considerable variability in the character of the leaf margins and degree of indument. The fruits of P. pubescens are reportedly gathered for food.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In Physalis hederifolia, the corolla limb is reflexed at maturity. This is a widespread species of the southwestern United States and the Great Plains (as far west as southern California and southeastern Nevada, and east into the western half of Iowa, Nebraska, Oklahoma, South Dakota, and Texas). Several varieties have been recognized based primarily on indument characters, but these features vary considerably over the range of the species. Plants from the more southern part of the range tend to have shorter hairs and to be more densely glandular; plants from northern Oklahoma and New Mexico northward tend to have longer hairs. All are clearly distinguishable from P. fendleri, which is eglandular, typically has forked or few-branched hairs, a distinctive leaf shape, and corollas with greenish-brownish smudges rather than distinct brown spots.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 14. FNA vol. 14.
Parent taxa Solanaceae > Physalis Solanaceae > Physalis
Sibling taxa
P. acutifolia, P. angulata, P. angustifolia, P. arenicola, P. caudella, P. cinerascens, P. cordata, P. crassifolia, P. fendleri, P. grisea, P. hederifolia, P. heterophylla, P. lanceolata, P. longifolia, P. missouriensis, P. mollis, P. neomexicana, P. philadelphica, P. pumila, P. solanacea, P. virginiana, P. walteri, P. ×elliottii
P. acutifolia, P. angulata, P. angustifolia, P. arenicola, P. caudella, P. cinerascens, P. cordata, P. crassifolia, P. fendleri, P. grisea, P. heterophylla, P. lanceolata, P. longifolia, P. missouriensis, P. mollis, P. neomexicana, P. philadelphica, P. pubescens, P. pumila, P. solanacea, P. virginiana, P. walteri, P. ×elliottii
Synonyms P. barbadensis, P. barbadensis var. glabra, P. floridana, P. latiphysa, P. pubescens var. glabra, P. pubescens var. integrifolia, P. turbinata P. comata, P. hederifolia var. comata, P. hederifolia var. palmeri, P. hederifolia var. puberula, P. palmeri, P. puberula, P. rotundata
Name authority Linnaeus: Sp. Pl. 1: 183. (1753) A. Gray: Proc. Amer. Acad. Arts 10: 65. (1874) — (as hederaefolia)
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