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common groundcherry, ground-cherry, long leafed tomatillo, long-leaf ground-cherry, wild tomatillo

ivy leafed tomatillo, ivyleaf groundcherry

Habit Herbs perennial, rhizomatous, rhizomes deeply buried, stout, glabrous or sparsely strigose, hairs simple, antrorse, to 0.5 mm. Herbs perennial, rhizomatous, rhizome stout, densely pubescent, hairs simple, sometimes jointed, glandular, 0.5–1 mm, sometimes also with sessile glands.
Stems

erect or erect to decumbent, branching frequently at distal nodes or several-branched from base, branches spreading or ascending, 1–6 dm.

erect to decumbent, usually branching from base and at most nodes, branches spreading, 0.5–3 dm.

Leaves

petiolate;

petiole 1/5–2/5 blade;

blade ovate to ovate-lanceolate or broadly lanceolate, 2.5–10(–13) × 0.5–6(–7) cm, base truncate to rounded, margins entire to coarsely dentate or irregularly crenate-dentate with only a few teeth.

petiolate;

petiole 1/2 to ± as long as blade;

blade broadly ovate to orbiculate, 1.5–3.5 × 1–3 cm, base cordate to rounded, margins ± entire or coarsely dentate, teeth sharp to blunt.

Pedicels

5–18 mm, 12–35 mm in fruit.

4–8(–13) mm, 5–15 mm in fruit.

Flowers

calyx (5–)7–12 mm, sparsely strigose with short, antrorse hairs, lobes 3–6 mm;

corolla yellow with 5 purple-brown smudges, campanulate-rotate, 10–20 mm;

anthers yellow or blue-tinged, not twisted after dehiscence, 2–4 mm.

calyx 5–7(–10) mm, lobes 1.5–3.5(–5) mm;

corolla yellow with 5 dark brown spots, campanulate-rotate, 7–12 mm;

anthers yellow, not twisted after dehiscence, 2–4 mm.

Fruiting calyces

loosely enclosing berry, 10-ribbed, 20–40 × 15–30 mm.

loosely enclosing berry, 10-ribbed, 20–30 × 15–25(–30) mm.

2n

= 24.

= 24.

Physalis longifolia

Physalis hederifolia

Phenology Flowering Apr–Aug.
Habitat Dry open gravelly sites, rocky ledges, open plains.
Elevation 200–2600 m. (700–8500 ft.)
Distribution
from USDA
North America; n Mexico [Introduced in Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; KS; MT; NE; NM; NV; OK; SD; TX; UT; WY; Mexico (Baja California, Chihuahua, Coahuila, Durango, Nuevo León, San Luis Potosí, Sonora, Tamaulipas, Zacatecas)
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In Physalis hederifolia, the corolla limb is reflexed at maturity. This is a widespread species of the southwestern United States and the Great Plains (as far west as southern California and southeastern Nevada, and east into the western half of Iowa, Nebraska, Oklahoma, South Dakota, and Texas). Several varieties have been recognized based primarily on indument characters, but these features vary considerably over the range of the species. Plants from the more southern part of the range tend to have shorter hairs and to be more densely glandular; plants from northern Oklahoma and New Mexico northward tend to have longer hairs. All are clearly distinguishable from P. fendleri, which is eglandular, typically has forked or few-branched hairs, a distinctive leaf shape, and corollas with greenish-brownish smudges rather than distinct brown spots.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems erect to decumbent, branching at base; leaf blades glabrous.
var. texana
1. Stems erect, branching at distal nodes; leaf blades glabrous or sparsely strigose.
→ 2
2. Leaf blades narrowly ovate-lanceolate to lan­ceolate; anthers yellow.
var. longifolia
2. Leaf blades ovate to ovate-lanceolate or broadly lanceolate; anthers blue-tinged.
var. subglabrata
Source FNA vol. 14. FNA vol. 14.
Parent taxa Solanaceae > Physalis Solanaceae > Physalis
Sibling taxa
P. acutifolia, P. angulata, P. angustifolia, P. arenicola, P. caudella, P. cinerascens, P. cordata, P. crassifolia, P. fendleri, P. grisea, P. hederifolia, P. heterophylla, P. lanceolata, P. missouriensis, P. mollis, P. neomexicana, P. philadelphica, P. pubescens, P. pumila, P. solanacea, P. virginiana, P. walteri, P. ×elliottii
P. acutifolia, P. angulata, P. angustifolia, P. arenicola, P. caudella, P. cinerascens, P. cordata, P. crassifolia, P. fendleri, P. grisea, P. heterophylla, P. lanceolata, P. longifolia, P. missouriensis, P. mollis, P. neomexicana, P. philadelphica, P. pubescens, P. pumila, P. solanacea, P. virginiana, P. walteri, P. ×elliottii
Subordinate taxa
P. longifolia var. longifolia, P. longifolia var. subglabrata, P. longifolia var. texana
Synonyms P. comata, P. hederifolia var. comata, P. hederifolia var. palmeri, P. hederifolia var. puberula, P. palmeri, P. puberula, P. rotundata
Name authority Nuttall: Trans. Amer. Philos. Soc., n. s. 5: 193. (1836) A. Gray: Proc. Amer. Acad. Arts 10: 65. (1874) — (as hederaefolia)
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