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thick leaf ground cherry, yellow nightshade groundcherry

clammy ground-cherry, ground cherry

Habit Herbs perennial, becoming suffrutescent, rhizomatous, rhizomes often just below soil surface, vertical, stout, puberulent, hairs divergent, to 0.5 mm, some glandular, appearing ± glabrous without magnification. Herbs perennial, rhizomatous, rhizomes deeply buried, stout, densely villous, hairs simple, jointed, divergent, 1–2 mm, sometimes also with shorter glandular hairs.
Stems

erect, branching from near base and at most nodes, branches widely spreading, distinctly zigzag, slender, 1–4(–10) dm.

erect to decumbent, branching at most nodes, branches spreading and decumbent, 1.5–10 dm.

Leaves

petiolate;

petiole mostly as long as blade;

blade broadly ovate to deltate, 0.8–3.3(–4.5) × 0.8–3.3(–4.5) cm, base cordate, sometimes slightly unequal, margins entire to unevenly coarsely dentate, sometimes thick and slightly succulent.

petiolate;

petiole 1/3–2/3 blade;

blade broadly ovate to suborbiculate, (2–)4–11(–13) × 3–9(–10) cm, base truncate to slightly cordate, margins deeply and irregularly dentate to ± entire.

Pedicels

8–24(–33) mm, (11–)14–30(–35) mm in fruit.

9–15(–20) mm, 20–30 mm in fruit.

Flowers

calyx (3–)4–7(–8) mm, lobes 1–3 mm;

corolla pale yellow with yellow or greenish-brown smudges or tinge, campanulate-rotate, 8–14 mm;

anthers yellow, not twisted after dehiscence, 1.5–3 mm.

calyx 6–12 mm, lobes 3–6 mm;

corolla yellow with 5 large purple-brown smudges, campanulate-rotate, 10–17 mm;

anthers yellow, rarely blue-tinged, not twisted after dehiscence, 2.5–4.5 mm.

Fruiting calyces

loosely enclosing berry, 10-ribbed, 14–30(–40) × (10–)15–20(–25) mm.

loosely enclosing berry, 10-ribbed, 25–40 × 15–30 mm.

2n

= 24.

= 24.

Physalis crassifolia

Physalis heterophylla

Phenology Flowering year-round in areas without frost, mostly Mar–Apr. Flowering May–Sep.
Habitat Gravelly or sandy slopes, washes, roadsides, mesas, canyons. Openings in hardwood forests, edges of pine woods, grasslands, fields, roadsides, dis­turbed sites.
Elevation 100–1700 m. (300–5600 ft.) 10–400 m. (0–1300 ft.)
Distribution
from FNA
AZ; CA; NV; UT; Mexico (Baja California, Baja California Sur, Chihuahua, Sinaloa, Sonora) [Introduced in Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; WY; ON; QC; SK
[WildflowerSearch map]
[BONAP county map]
Discussion

In Physalis crassifolia, the corolla limb is widely flaring and reflexed when the flower is fully open, and the flowers are more nodding than fully pendent. Some herbarium specimen labels indicate that plants flower the first year. Physalis greenei Vasey & Rose, not validly published, has been misapplied to some representatives of P. crassifolia. Physalis crassifolia is widespread in Arizona, but it is restricted in California to south-southeastern counties as far north as Inyo, in Nevada to Clark and Lincoln counties, and in Utah to Washington County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Physalis heterophylla is widespread east of the Rocky Mountains. Herbarium specimens from Manitoba, Oregon, and Utah represent historical collections. Morphological variation has been recognized taxonomically in some manuals, although intergradation occurs among varieties and they often cannot be identified reliably. The cultivated P. peruviana (cape gooseberry) is similar to P. heterophylla except that it is not glandular and has shorter pedicels (6–8 mm in flower, 13–15 mm in fruit). W. F. Hinton (1975b) reported the uncommon occurrence of a population in North Carolina representing natural hybridization between P. heterophylla and P. virginiana.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 14. FNA vol. 14.
Parent taxa Solanaceae > Physalis Solanaceae > Physalis
Sibling taxa
P. acutifolia, P. angulata, P. angustifolia, P. arenicola, P. caudella, P. cinerascens, P. cordata, P. fendleri, P. grisea, P. hederifolia, P. heterophylla, P. lanceolata, P. longifolia, P. missouriensis, P. mollis, P. neomexicana, P. philadelphica, P. pubescens, P. pumila, P. solanacea, P. virginiana, P. walteri, P. ×elliottii
P. acutifolia, P. angulata, P. angustifolia, P. arenicola, P. caudella, P. cinerascens, P. cordata, P. crassifolia, P. fendleri, P. grisea, P. hederifolia, P. lanceolata, P. longifolia, P. missouriensis, P. mollis, P. neomexicana, P. philadelphica, P. pubescens, P. pumila, P. solanacea, P. virginiana, P. walteri, P. ×elliottii
Synonyms P. crassifolia var. cardiophylla, P. crassifolia var. versicolor, P. versicolor P. ambigua, P. heterophylla var. ambigua, P. heterophylla var. clavipes, P. heterophylla var. nyctaginea, P. heterophylla var. villosa, P. nyctaginea, P. sinuata
Name authority Bentham: Bot. Voy. Sulphur, 40. (1844) Nees: Linnaea 6: 463. (1831)
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