Physalis crassifolia |
Physalis heterophylla |
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thick leaf ground cherry, yellow nightshade groundcherry |
clammy ground-cherry, ground cherry |
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Habit | Herbs perennial, becoming suffrutescent, rhizomatous, rhizomes often just below soil surface, vertical, stout, puberulent, hairs divergent, to 0.5 mm, some glandular, appearing ± glabrous without magnification. | Herbs perennial, rhizomatous, rhizomes deeply buried, stout, densely villous, hairs simple, jointed, divergent, 1–2 mm, sometimes also with shorter glandular hairs. |
Stems | erect, branching from near base and at most nodes, branches widely spreading, distinctly zigzag, slender, 1–4(–10) dm. |
erect to decumbent, branching at most nodes, branches spreading and decumbent, 1.5–10 dm. |
Leaves | petiolate; petiole mostly as long as blade; blade broadly ovate to deltate, 0.8–3.3(–4.5) × 0.8–3.3(–4.5) cm, base cordate, sometimes slightly unequal, margins entire to unevenly coarsely dentate, sometimes thick and slightly succulent. |
petiolate; petiole 1/3–2/3 blade; blade broadly ovate to suborbiculate, (2–)4–11(–13) × 3–9(–10) cm, base truncate to slightly cordate, margins deeply and irregularly dentate to ± entire. |
Pedicels | 8–24(–33) mm, (11–)14–30(–35) mm in fruit. |
9–15(–20) mm, 20–30 mm in fruit. |
Flowers | calyx (3–)4–7(–8) mm, lobes 1–3 mm; corolla pale yellow with yellow or greenish-brown smudges or tinge, campanulate-rotate, 8–14 mm; anthers yellow, not twisted after dehiscence, 1.5–3 mm. |
calyx 6–12 mm, lobes 3–6 mm; corolla yellow with 5 large purple-brown smudges, campanulate-rotate, 10–17 mm; anthers yellow, rarely blue-tinged, not twisted after dehiscence, 2.5–4.5 mm. |
Fruiting calyces | loosely enclosing berry, 10-ribbed, 14–30(–40) × (10–)15–20(–25) mm. |
loosely enclosing berry, 10-ribbed, 25–40 × 15–30 mm. |
2n | = 24. |
= 24. |
Physalis crassifolia |
Physalis heterophylla |
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Phenology | Flowering year-round in areas without frost, mostly Mar–Apr. | Flowering May–Sep. |
Habitat | Gravelly or sandy slopes, washes, roadsides, mesas, canyons. | Openings in hardwood forests, edges of pine woods, grasslands, fields, roadsides, disturbed sites. |
Elevation | 100–1700 m. (300–5600 ft.) | 10–400 m. (0–1300 ft.) |
Distribution |
AZ; CA; NV; UT; Mexico (Baja California, Baja California Sur, Chihuahua, Sinaloa, Sonora) [Introduced in Australia]
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AL; AR; CO; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; ND; NE; NH; NJ; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; WY; ON; QC; SK
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Discussion | In Physalis crassifolia, the corolla limb is widely flaring and reflexed when the flower is fully open, and the flowers are more nodding than fully pendent. Some herbarium specimen labels indicate that plants flower the first year. Physalis greenei Vasey & Rose, not validly published, has been misapplied to some representatives of P. crassifolia. Physalis crassifolia is widespread in Arizona, but it is restricted in California to south-southeastern counties as far north as Inyo, in Nevada to Clark and Lincoln counties, and in Utah to Washington County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Physalis heterophylla is widespread east of the Rocky Mountains. Herbarium specimens from Manitoba, Oregon, and Utah represent historical collections. Morphological variation has been recognized taxonomically in some manuals, although intergradation occurs among varieties and they often cannot be identified reliably. The cultivated P. peruviana (cape gooseberry) is similar to P. heterophylla except that it is not glandular and has shorter pedicels (6–8 mm in flower, 13–15 mm in fruit). W. F. Hinton (1975b) reported the uncommon occurrence of a population in North Carolina representing natural hybridization between P. heterophylla and P. virginiana. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 14. | FNA vol. 14. |
Parent taxa | Solanaceae > Physalis | Solanaceae > Physalis |
Sibling taxa | ||
Synonyms | P. crassifolia var. cardiophylla, P. crassifolia var. versicolor, P. versicolor | P. ambigua, P. heterophylla var. ambigua, P. heterophylla var. clavipes, P. heterophylla var. nyctaginea, P. heterophylla var. villosa, P. nyctaginea, P. sinuata |
Name authority | Bentham: Bot. Voy. Sulphur, 40. (1844) | Nees: Linnaea 6: 463. (1831) |
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