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thick leaf ground cherry, yellow nightshade groundcherry

ivy leafed tomatillo, ivyleaf groundcherry

Habit Herbs perennial, becoming suffrutescent, rhizomatous, rhizomes often just below soil surface, vertical, stout, puberulent, hairs divergent, to 0.5 mm, some glandular, appearing ± glabrous without magnification. Herbs perennial, rhizomatous, rhizome stout, densely pubescent, hairs simple, sometimes jointed, glandular, 0.5–1 mm, sometimes also with sessile glands.
Stems

erect, branching from near base and at most nodes, branches widely spreading, distinctly zigzag, slender, 1–4(–10) dm.

erect to decumbent, usually branching from base and at most nodes, branches spreading, 0.5–3 dm.

Leaves

petiolate;

petiole mostly as long as blade;

blade broadly ovate to deltate, 0.8–3.3(–4.5) × 0.8–3.3(–4.5) cm, base cordate, sometimes slightly unequal, margins entire to unevenly coarsely dentate, sometimes thick and slightly succulent.

petiolate;

petiole 1/2 to ± as long as blade;

blade broadly ovate to orbiculate, 1.5–3.5 × 1–3 cm, base cordate to rounded, margins ± entire or coarsely dentate, teeth sharp to blunt.

Pedicels

8–24(–33) mm, (11–)14–30(–35) mm in fruit.

4–8(–13) mm, 5–15 mm in fruit.

Flowers

calyx (3–)4–7(–8) mm, lobes 1–3 mm;

corolla pale yellow with yellow or greenish-brown smudges or tinge, campanulate-rotate, 8–14 mm;

anthers yellow, not twisted after dehiscence, 1.5–3 mm.

calyx 5–7(–10) mm, lobes 1.5–3.5(–5) mm;

corolla yellow with 5 dark brown spots, campanulate-rotate, 7–12 mm;

anthers yellow, not twisted after dehiscence, 2–4 mm.

Fruiting calyces

loosely enclosing berry, 10-ribbed, 14–30(–40) × (10–)15–20(–25) mm.

loosely enclosing berry, 10-ribbed, 20–30 × 15–25(–30) mm.

2n

= 24.

= 24.

Physalis crassifolia

Physalis hederifolia

Phenology Flowering year-round in areas without frost, mostly Mar–Apr. Flowering Apr–Aug.
Habitat Gravelly or sandy slopes, washes, roadsides, mesas, canyons. Dry open gravelly sites, rocky ledges, open plains.
Elevation 100–1700 m. (300–5600 ft.) 200–2600 m. (700–8500 ft.)
Distribution
from FNA
AZ; CA; NV; UT; Mexico (Baja California, Baja California Sur, Chihuahua, Sinaloa, Sonora) [Introduced in Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; KS; MT; NE; NM; NV; OK; SD; TX; UT; WY; Mexico (Baja California, Chihuahua, Coahuila, Durango, Nuevo León, San Luis Potosí, Sonora, Tamaulipas, Zacatecas)
[WildflowerSearch map]
[BONAP county map]
Discussion

In Physalis crassifolia, the corolla limb is widely flaring and reflexed when the flower is fully open, and the flowers are more nodding than fully pendent. Some herbarium specimen labels indicate that plants flower the first year. Physalis greenei Vasey & Rose, not validly published, has been misapplied to some representatives of P. crassifolia. Physalis crassifolia is widespread in Arizona, but it is restricted in California to south-southeastern counties as far north as Inyo, in Nevada to Clark and Lincoln counties, and in Utah to Washington County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In Physalis hederifolia, the corolla limb is reflexed at maturity. This is a widespread species of the southwestern United States and the Great Plains (as far west as southern California and southeastern Nevada, and east into the western half of Iowa, Nebraska, Oklahoma, South Dakota, and Texas). Several varieties have been recognized based primarily on indument characters, but these features vary considerably over the range of the species. Plants from the more southern part of the range tend to have shorter hairs and to be more densely glandular; plants from northern Oklahoma and New Mexico northward tend to have longer hairs. All are clearly distinguishable from P. fendleri, which is eglandular, typically has forked or few-branched hairs, a distinctive leaf shape, and corollas with greenish-brownish smudges rather than distinct brown spots.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 14. FNA vol. 14.
Parent taxa Solanaceae > Physalis Solanaceae > Physalis
Sibling taxa
P. acutifolia, P. angulata, P. angustifolia, P. arenicola, P. caudella, P. cinerascens, P. cordata, P. fendleri, P. grisea, P. hederifolia, P. heterophylla, P. lanceolata, P. longifolia, P. missouriensis, P. mollis, P. neomexicana, P. philadelphica, P. pubescens, P. pumila, P. solanacea, P. virginiana, P. walteri, P. ×elliottii
P. acutifolia, P. angulata, P. angustifolia, P. arenicola, P. caudella, P. cinerascens, P. cordata, P. crassifolia, P. fendleri, P. grisea, P. heterophylla, P. lanceolata, P. longifolia, P. missouriensis, P. mollis, P. neomexicana, P. philadelphica, P. pubescens, P. pumila, P. solanacea, P. virginiana, P. walteri, P. ×elliottii
Synonyms P. crassifolia var. cardiophylla, P. crassifolia var. versicolor, P. versicolor P. comata, P. hederifolia var. comata, P. hederifolia var. palmeri, P. hederifolia var. puberula, P. palmeri, P. puberula, P. rotundata
Name authority Bentham: Bot. Voy. Sulphur, 40. (1844) A. Gray: Proc. Amer. Acad. Arts 10: 65. (1874) — (as hederaefolia)
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