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thick leaf ground cherry, yellow nightshade groundcherry

Habit Herbs perennial, becoming suffrutescent, rhizomatous, rhizomes often just below soil surface, vertical, stout, puberulent, hairs divergent, to 0.5 mm, some glandular, appearing ± glabrous without magnification. Herbs perennial, rhizomatous, rhizome stout, pubescent, hairs simple, forked, or 3-branched, to 0.5 mm, most branching at base and branches appressed to surface, giving plants a grayish appearance.
Stems

erect, branching from near base and at most nodes, branches widely spreading, distinctly zigzag, slender, 1–4(–10) dm.

erect, usually branching from base and most nodes, branches spreading, 0.5–5 dm.

Leaves

petiolate;

petiole mostly as long as blade;

blade broadly ovate to deltate, 0.8–3.3(–4.5) × 0.8–3.3(–4.5) cm, base cordate, sometimes slightly unequal, margins entire to unevenly coarsely dentate, sometimes thick and slightly succulent.

petiolate;

petiole 1/3–2/3 blade;

blade ovate-lanceolate to ovate, 1–5.5 × 1–3.5 cm, base deltate-truncate to slightly cordate, sometimes unequal, margins sinuate to coarsely and irregularly dentate, teeth few, sometimes only 1 tooth per side near base.

Pedicels

8–24(–33) mm, (11–)14–30(–35) mm in fruit.

3–11(–17) mm, 10–15(–20) mm in fruit.

Flowers

calyx (3–)4–7(–8) mm, lobes 1–3 mm;

corolla pale yellow with yellow or greenish-brown smudges or tinge, campanulate-rotate, 8–14 mm;

anthers yellow, not twisted after dehiscence, 1.5–3 mm.

calyx 5–8 mm, lobes 2–4 mm;

corolla yellow with 5 greenish to brown smudges, campanulate-rotate, (7–)10–12 mm;

anthers yellow or purple-tinged, not twisted after dehiscence, 1.5–3 mm.

Fruiting calyces

loosely enclosing berry, 10-ribbed, 14–30(–40) × (10–)15–20(–25) mm.

loosely enclosing berry, 10-ribbed, 17–30(–35) × 15–25 mm.

2n

= 24.

= 24.

Physalis crassifolia

Physalis fendleri

Phenology Flowering year-round in areas without frost, mostly Mar–Apr. Flowering May–Sep.
Habitat Gravelly or sandy slopes, washes, roadsides, mesas, canyons. Rocky to sandy soil, loose slopes, pinyon-juniper-ponderosa zones.
Elevation 100–1700 m. (300–5600 ft.) 1300–2300 m. (4300–7500 ft.)
Distribution
from FNA
AZ; CA; NV; UT; Mexico (Baja California, Baja California Sur, Chihuahua, Sinaloa, Sonora) [Introduced in Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; NM; NV; OK; TX; UT; Mexico (Baja California, San Luis Potosí, Tamaulipas)
Discussion

In Physalis crassifolia, the corolla limb is widely flaring and reflexed when the flower is fully open, and the flowers are more nodding than fully pendent. Some herbarium specimen labels indicate that plants flower the first year. Physalis greenei Vasey & Rose, not validly published, has been misapplied to some representatives of P. crassifolia. Physalis crassifolia is widespread in Arizona, but it is restricted in California to south-southeastern counties as far north as Inyo, in Nevada to Clark and Lincoln counties, and in Utah to Washington County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In Physalis fendleri, the corolla limb is reflexed at maturity. Some populations in New Mexico have mostly retrorse, and very few branched, hairs. In the flora area, P. fendleri can be found as far west as southern California, northward into southern Nevada, Utah, and Colorado, and eastward into western Oklahoma and Texas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 14. FNA vol. 14.
Parent taxa Solanaceae > Physalis Solanaceae > Physalis
Sibling taxa
P. acutifolia, P. angulata, P. angustifolia, P. arenicola, P. caudella, P. cinerascens, P. cordata, P. fendleri, P. grisea, P. hederifolia, P. heterophylla, P. lanceolata, P. longifolia, P. missouriensis, P. mollis, P. neomexicana, P. philadelphica, P. pubescens, P. pumila, P. solanacea, P. virginiana, P. walteri, P. ×elliottii
P. acutifolia, P. angulata, P. angustifolia, P. arenicola, P. caudella, P. cinerascens, P. cordata, P. crassifolia, P. grisea, P. hederifolia, P. heterophylla, P. lanceolata, P. longifolia, P. missouriensis, P. mollis, P. neomexicana, P. philadelphica, P. pubescens, P. pumila, P. solanacea, P. virginiana, P. walteri, P. ×elliottii
Synonyms P. crassifolia var. cardiophylla, P. crassifolia var. versicolor, P. versicolor P. fendleri var. cordifolia, P. hederifolia var. cordifolia, P. hederifolia var. fendleri
Name authority Bentham: Bot. Voy. Sulphur, 40. (1844) A. Gray: Proc. Amer. Acad. Arts 10: 66. (1874)
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