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gale of the wind

gulf leaf-flower

Habit Herbs, annual, monoecious, 1–5 dm; branching phyllanthoid. Herbs, annual, monoecious, 1–4 dm; branching phyllanthoid.
Stems

main stems terete, not winged, glabrous; ultimate branchlets subterete, not winged, glabrous.

main stems angled, not winged, glabrous; ultimate branchlets subterete, narrowly winged, ± scabridulous.

Leaves

on main stems spiral, scalelike;

stipules not auriculate, brown.;

leaves on ultimate branchlets distichous, well developed;

stipules not auriculate, brown;

blade elliptic, 11–20 × 4.5–9 mm, base obtuse to rounded, apex obtuse, both surfaces glabrous.

on main stems spiral, scalelike;

stipules not auriculate, pale green to nearly white.;

leaves on ultimate branchlets distichous, well developed;

stipules not auriculate, pale green to nearly white;

blade elliptic-oblong, 6–11 × 3–5 mm, base cuneate to obtuse, apex rounded, both surfaces glabrous.

Inflorescences

cymules or flowers solitary, unisexual, proximal with 3–7 staminate flowers, distal with 1 pistillate flower.

cymules or flowers solitary, unisexual, proximal with 2–3 staminate flowers, distal with 1 pistillate flower.

Pedicels

staminate 1.2–1.8 mm, pistillate spreading in fruit, 4–7 mm.

staminate 0.2–0.5 mm, pistillate spreading in fruit, 1.3–2 mm.

Staminate flowers

sepals 5(–6), pale green, flat, 1.5–3 mm;

nectary extrastaminal, 5(–6) glands;

stamens 3, filaments connate 1/2 length.

sepals 6, white to pale yellow, flat, 0.4–0.7 mm;

nectary extrastaminal, 6 glands;

stamens 3, filaments connate throughout.

Pistillate flowers

sepals 5, green, flat, 3–3.5 mm, pinnately veined;

nectary annular, unlobed.

sepals 6, green with broad white margins, flat, (1–)1.2–1.5 mm, 1-veined;

nectary annular, 6–9-lobed.

Capsules

3.5 mm diam., smooth.

2.1 mm diam., smooth.

Seeds

uniformly brown, 1.5–1.8 mm, verrucose.

uniformly brown, 0.9–1.1 mm, longitudinally ribbed.

2n

= 26 (Costa Rica).

Phyllanthus niruri

Phyllanthus fraternus

Phenology Flowering and fruiting late summer–fall. Flowering and fruiting year-round.
Habitat River and stream banks, sand. Fields, roadsides, gardens, other disturbed areas.
Elevation 60–120 m. (200–400 ft.) 0–30 m. (0–100 ft.)
Distribution
from USDA
Mexico; Central America; South America; West Indies
[BONAP county map]
from FNA
FL; GA; LA; MS; SC; TX; s Asia (India, Pakistan) [Introduced in North America; introduced also in West Indies, Bermuda, w Asia, Africa, Indian Ocean Islands]
[BONAP county map]
Discussion

Phyllanthus niruri is found in the flora area only in DeWitt, Fayette, and Lavaca counties (and historically from Gonzales County, where it appears to be extirpated; L. E. Brown and S. J. Marcus 1998); it is widespread in the American tropics. Like P. urinaria, it is widely used in folk medicine and is the subject of intense pharmacological research. Plants from outside the West Indies and Caribbean northern South America often have been segregated as subsp. lathyroides; the differences are trivial and recent authors (G. L. Webster 2001; V. W. Steinmann 2007) did not subdivide the species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Phyllanthus fraternus, native to Pakistan and northwest India, apparently was introduced into the United States in the 1950s, first in Louisiana, and has spread mainly along the Gulf Coast.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 342. FNA vol. 12, p. 342.
Parent taxa Phyllanthaceae > Phyllanthus Phyllanthaceae > Phyllanthus
Sibling taxa
P. abnormis, P. acidus, P. amarus, P. angustifolius, P. caroliniensis, P. ericoides, P. evanescens, P. fluitans, P. fraternus, P. liebmannianus, P. pentaphyllus, P. polygonoides, P. tenellus, P. urinaria, P. warnockii
P. abnormis, P. acidus, P. amarus, P. angustifolius, P. caroliniensis, P. ericoides, P. evanescens, P. fluitans, P. liebmannianus, P. niruri, P. pentaphyllus, P. polygonoides, P. tenellus, P. urinaria, P. warnockii
Synonyms P. lathyroides, P. niruri subsp. lathyroides P. niruri var. scabrellus
Name authority Linnaeus: Sp. Pl. 2: 981. (1753) G. L. Webster: Contr. Gray Herb. 176: 53. (1955)
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