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abnormal leaflower, Drummond's leaf-flower

gale of the wind

Habit Herbs, annual, sometimes becoming woody and then appearing to be perennial, monoecious, 1–5 dm; branching phyllanthoid. Herbs, annual, monoecious, 1–5 dm; branching phyllanthoid.
Stems

main stems persistent, terete, not winged, glabrous or scabridulous; ultimate branchlets deciduous, subterete, not winged, glabrous or scabridulous.

main stems terete, not winged, glabrous; ultimate branchlets subterete, not winged, glabrous.

Leaves

on main stems spiral, scalelike;

stipules not auriculate, pale reddish brown.;

leaves on ultimate branchlets distichous, well developed;

stipules not auriculate, greenish white;

blade elliptic to oblong, 3–10 × 1–4 mm, base cuneate to subcordate, apex obtuse to emarginate, both surfaces glabrous or scabridulous.

on main stems spiral, scalelike;

stipules not auriculate, brown.;

leaves on ultimate branchlets distichous, well developed;

stipules not auriculate, brown;

blade elliptic, 11–20 × 4.5–9 mm, base obtuse to rounded, apex obtuse, both surfaces glabrous.

Inflorescences

cymules, proximal staminate with 2 flowers, distal bisexual with 1 pistillate flower and 1–3 staminate flowers.

cymules or flowers solitary, unisexual, proximal with 3–7 staminate flowers, distal with 1 pistillate flower.

Pedicels

staminate 0.7–1.5 mm, pistillate spreading in fruit, (1–)1.5–3(–3.5) mm.

staminate 1.2–1.8 mm, pistillate spreading in fruit, 4–7 mm.

Staminate flowers

sepals 5–6 in basal cymules, 4 in distal cymules, pale yellowish green, sometimes suffused with red, flat, 0.5–1 mm;

nectary extrastaminal, 4(–6) glands;

stamens 2(–3 in basal cymules), filaments connate throughout.

sepals 5(–6), pale green, flat, 1.5–3 mm;

nectary extrastaminal, 5(–6) glands;

stamens 3, filaments connate 1/2 length.

Pistillate flowers

sepals 5–6, green with nearly white margins, flat, (0.5–)0.7–1.1 mm, 1-veined;

nectary 3 glands (2-fid distally, thus resembling 6 glands).

sepals 5, green, flat, 3–3.5 mm, pinnately veined;

nectary annular, unlobed.

Capsules

2.3–2.7 mm diam., smooth.

3.5 mm diam., smooth.

Seeds

uniformly brown, 1.1–1.5 mm, longitudinally ribbed.

uniformly brown, 1.5–1.8 mm, verrucose.

2n

= 26 (Costa Rica).

Phyllanthus abnormis

Phyllanthus niruri

Phenology Flowering and fruiting late summer–fall.
Habitat River and stream banks, sand.
Elevation 60–120 m. (200–400 ft.)
Distribution
from FNA
FL; NM; OK; TX; ne Mexico
[WildflowerSearch map]
[BONAP county map]
from USDA
Mexico; Central America; South America; West Indies
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Phyllanthus niruri is found in the flora area only in DeWitt, Fayette, and Lavaca counties (and historically from Gonzales County, where it appears to be extirpated; L. E. Brown and S. J. Marcus 1998); it is widespread in the American tropics. Like P. urinaria, it is widely used in folk medicine and is the subject of intense pharmacological research. Plants from outside the West Indies and Caribbean northern South America often have been segregated as subsp. lathyroides; the differences are trivial and recent authors (G. L. Webster 2001; V. W. Steinmann 2007) did not subdivide the species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Pistillate nectary glands strongly unequal, spatulate, as long as or longer than broad; leaf blades glabrous on both surfaces or sparsely to moderately scabridulous abaxially; bisexual cymules with 1 staminate flower.
var. abnormis
1. Pistillate nectary glands subequal, reniform, broader than long; leaf blades densely scabridulous on both surfaces; bisexual cymules with 1–3 staminate flowers.
var. riograndensis
Source FNA vol. 12, p. 343. FNA vol. 12, p. 342.
Parent taxa Phyllanthaceae > Phyllanthus Phyllanthaceae > Phyllanthus
Sibling taxa
P. acidus, P. amarus, P. angustifolius, P. caroliniensis, P. ericoides, P. evanescens, P. fluitans, P. fraternus, P. liebmannianus, P. niruri, P. pentaphyllus, P. polygonoides, P. tenellus, P. urinaria, P. warnockii
P. abnormis, P. acidus, P. amarus, P. angustifolius, P. caroliniensis, P. ericoides, P. evanescens, P. fluitans, P. fraternus, P. liebmannianus, P. pentaphyllus, P. polygonoides, P. tenellus, P. urinaria, P. warnockii
Subordinate taxa
P. abnormis var. abnormis, P. abnormis var. riograndensis
Synonyms P. lathyroides, P. niruri subsp. lathyroides
Name authority Baillon: Adansonia 1: 42. (1860) Linnaeus: Sp. Pl. 2: 981. (1753)
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