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abnormal leaflower, Drummond's leaf-flower

heather leaf-flower

Habit Herbs, annual, sometimes becoming woody and then appearing to be perennial, monoecious, 1–5 dm; branching phyllanthoid. Shrubs, monoecious, 0.5–2 dm; branching not phyllanthoid.
Stems

main stems persistent, terete, not winged, glabrous or scabridulous; ultimate branchlets deciduous, subterete, not winged, glabrous or scabridulous.

terete, not winged, glabrous.

Leaves

on main stems spiral, scalelike;

stipules not auriculate, pale reddish brown.;

leaves on ultimate branchlets distichous, well developed;

stipules not auriculate, greenish white;

blade elliptic to oblong, 3–10 × 1–4 mm, base cuneate to subcordate, apex obtuse to emarginate, both surfaces glabrous or scabridulous.

persistent, spiral, all well developed;

stipules auriculate, pink with narrow white margins;

blade narrowly oblanceolate, 1.5–3.5(–5) × 0.7–1.5 mm, base obtuse to rounded, apex pungent, both surfaces glabrous.

Inflorescences

cymules, proximal staminate with 2 flowers, distal bisexual with 1 pistillate flower and 1–3 staminate flowers.

solitary flowers, staminate proximal, pistillate distal.

Pedicels

staminate 0.7–1.5 mm, pistillate spreading in fruit, (1–)1.5–3(–3.5) mm.

staminate 0.6–0.8 mm, pistillate spreading in fruit, 0.9–1.2 mm.

Staminate flowers

sepals 5–6 in basal cymules, 4 in distal cymules, pale yellowish green, sometimes suffused with red, flat, 0.5–1 mm;

nectary extrastaminal, 4(–6) glands;

stamens 2(–3 in basal cymules), filaments connate throughout.

sepals 6, pale brownish green, flat, 0.9–1.2 mm;

nectary extrastaminal, 6 glands;

stamens 3, filaments connate throughout.

Pistillate flowers

sepals 5–6, green with nearly white margins, flat, (0.5–)0.7–1.1 mm, 1-veined;

nectary 3 glands (2-fid distally, thus resembling 6 glands).

sepals 6, pale brownish green, flat, 1.1–1.4 mm, obscurely veined;

nectary annular, 6-lobed.

Capsules

2.3–2.7 mm diam., smooth.

2 mm diam., smooth.

Seeds

uniformly brown, 1.1–1.5 mm, longitudinally ribbed.

uniformly brown, 0.9–1 mm, verrucose.

Phyllanthus abnormis

Phyllanthus ericoides

Phenology Flowering and fruiting year-round, following rains.
Habitat Desert scrublands on limestone rocks.
Elevation 600–700 m. (2000–2300 ft.)
Distribution
from FNA
FL; NM; OK; TX; ne Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico (Chihuahua, Coahuila)
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Phyllanthus ericoides is known from only a few populations in Brewster and Terrell counties, Texas, and nearby Chihuahua and Coahuila, Mexico. Although stunted plants of the related P. polygonoides may resemble P. ericoides, the latter may be distinguished by its woody habit, pungent leaves, shorter pedicels, completely connate filaments, and smaller seeds. In addition, the woody base of P. ericoides is reported to smell like coffee with chicory (M. C. Johnston and B. H. Warnock 1963).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Pistillate nectary glands strongly unequal, spatulate, as long as or longer than broad; leaf blades glabrous on both surfaces or sparsely to moderately scabridulous abaxially; bisexual cymules with 1 staminate flower.
var. abnormis
1. Pistillate nectary glands subequal, reniform, broader than long; leaf blades densely scabridulous on both surfaces; bisexual cymules with 1–3 staminate flowers.
var. riograndensis
Source FNA vol. 12, p. 343. FNA vol. 12, p. 338.
Parent taxa Phyllanthaceae > Phyllanthus Phyllanthaceae > Phyllanthus
Sibling taxa
P. acidus, P. amarus, P. angustifolius, P. caroliniensis, P. ericoides, P. evanescens, P. fluitans, P. fraternus, P. liebmannianus, P. niruri, P. pentaphyllus, P. polygonoides, P. tenellus, P. urinaria, P. warnockii
P. abnormis, P. acidus, P. amarus, P. angustifolius, P. caroliniensis, P. evanescens, P. fluitans, P. fraternus, P. liebmannianus, P. niruri, P. pentaphyllus, P. polygonoides, P. tenellus, P. urinaria, P. warnockii
Subordinate taxa
P. abnormis var. abnormis, P. abnormis var. riograndensis
Name authority Baillon: Adansonia 1: 42. (1860) Torrey: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 193. (1859)
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