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cliff mock orange, streambank mock orange

Lewis' mock-orange, mock-orange, wild mock orange

Habit Shrubs, 5–20 dm. Shrubs, 15–40 dm.
Stems

brown, gray, or stramineous, branched, glabrous to densely strigose;

bark deciduous, exfoliating or flaking, reddish;

branches sprawling;

axillary buds exposed.

erect to ascending, green, weathering brown, gray, or stramineous, glabrous or sparsely strigose (especially at nodes);

bark deciduous, exfoliating or flaking, reddish;

axillary buds hidden in pouches, sometimes apex exposed, especially on vigorous sprout-shoots.

Leaves

petiole 1–9 mm;

blade white or gray abaxially, green adaxially, broadly lanceolate to broadly ovate or narrowly to broadly elliptic, 2–8 × 1–5.5 cm, herbaceous, base narrowly cuneate to rounded, margins entire or irregularly to regularly serrate, crenate, or dentate, plane, abaxial surface slightly to densely loosely strigose, hairs often gently curved or arched-twisted, 0.7–1.5 mm, adaxial surface glabrate to moderately strigose, hairs usually evenly distributed, when sparse, sometimes distributed mainly along major veins, 0.2–0.8 mm, either thick-based and tightly antrorsely appressed or less thick-based, longer and looser, similar to abaxial hairs.

petiole 1–6 mm;

blade broadly lanceolate to broadly ovate, or narrowly to broadly elliptic, 1.5–10 × 1–5 cm, larger blades usually less than 6 × 2.5 cm, base narrowly cuneate to rounded or cordate, margins entire or irregularly to regularly serrate, crenate, or dentate, plane or slightly revolute, abaxial surface glabrous or sparsely strigose, hairs usually appressed-ascending, not twisted, main vein axils often densely strigose-tomentose, main veins sometimes sparsely strigose, secondary and tertiary veins rarely sparsely strigose, adaxial surface glabrous or sparsely to moderately strigose, especially near base and margins.

Inflorescences

cymes or cymose racemes, or flowers solitary, 1–3(–5)-flowered, proximal 2(–4) flowers often in axils of leaves or bracts.

usually cymose racemes or cymose panicles, sometimes flowers solitary, (1–)7–49-flowered, proximal 2, 4, or 6 flowers often in axils of nearly normal to much reduced (bracteal) leaves.

Pedicels

2–11 mm, moderately to densely strigose.

3–8 mm, glabrous or moderately strigose.

Flowers

hypanthium glabrous or densely strigose, hairs tightly or loosely appressed;

sepals ovate-lanceolate or triangular, 3–6 × 2–4 mm, apex acuminate, abaxial surface sparsely to densely strigose, adaxial surface glabrous except densely villous distally;

petals white, oblong, obovate, or orbiculate, 5–13 × 4–13 mm;

stamens 14–35;

filaments distinct, 3–9 mm;

anthers 1 × 0.7 mm;

style 1, clavate, 4–6 mm;

stigmatic portion 2–3 mm.

hypanthium glabrous or sparsely strigose, hairs scattered or concentrated on veins;

sepals ovate, ovate-lanceolate, or triangular, 5–8 × 3–5 mm, apex acute to acuminate, abaxial surface glabrous or sparsely strigose, adaxial surface glabrous except densely villous distally;

petals white, oblong, obovate, or orbiculate, 5–20(–25) × 4–15 mm;

stamens 25–40(–50);

filaments distinct, 5–11 mm;

anthers 2 × 1.5 mm;

styles 4, cylindric, 4–8 mm, connate proximally, lobes 1–4 × 0.4–0.5 mm;

stigmatic surfaces 1–3.5 mm.

Capsules

obconic to obovoid, (3–)4–7 × 3–6 mm.

obconic to obovoid, 7–11 × 5–7 mm.

Seeds

not caudate, 0.9–1.2 mm.

caudate, 3 mm.

2n

= 26.

= 26.

Philadelphus hirsutus

Philadelphus lewisii

Phenology Flowering Apr–May; fruiting Jun–Aug. Flowering May–Jul; fruiting Jun–Sep.
Habitat Bluffs, rock outcrops, seepage areas over rock, stream banks, particularly over calcareous sedimentary or mafic metamorphic or igneous rocks. Cliffs, rock outcrops, slopes in pine woodlands and forests, stream banks, talus, seasonally dry ravines.
Elevation 100–800 m. (300–2600 ft.) 0–2500 m. (0–8200 ft.)
Distribution
from FNA
AL; AR; GA; KY; MD; MS; NC; SC; TN; VA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; MT; OR; WA; AB; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

The occurrence of Philadelphus hirsutus in Maryland is believed to be due to an introduction. Philadelphus hirsutus is the most xerophytic of the native southeastern species of Philadelphus, often occurring on rock outcrops with only seasonal moisture. It is sometimes confused with P. pubescens; the undivided style and exposed buds are diagnostic.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Philadelphus lewisii, the state flower of Idaho, is rarely cultivated beyond its native range; there is no evidence that it is naturalized in other parts of North America.

A broadly defined Philadelphus lewisii is here recognized with some reluctance; it includes all previously named northwestern entities of subg. Philadelphus. Hu S. Y. (1954–1956) recognized 16 entities at varietal and specific rank, arrayed in two subgenera. C. L. Hitchcock et al. (1955–1969, vol. 3) described P. lewisii as being extremely variable in both vegetative and floral characters and further stated that most ecological or geographic races do not merit taxonomic recognition. Most recent floristic treatments have followed Hitchcock et al. or in some cases recognized two or three entities. The treatment of Hitchcock et al. is followed here, and there does not appear to be a case for the recognition of the two possible exceptions they mentioned (vars. gordonianus and parvifolius). Philadelphus californicus, P. cordifolius, and P. insignis were separated by Hu from P. lewisii at a subgeneric level based primarily on exposed buds and many-flowered inflorescences. The exposed buds are seen sporadically in P. lewisii, including outside the distribution accorded P. californicus, and are not evolutionarily cognate with the exposed buds of subg. Deutzioides; based on examination of herbarium material, they seem to be an expression of extra-vigorous growth, but additional study of plants in the field is needed. Some plants from the Sierra Nevada in California, considered by Hu to be occupied solely by P. californicus, are indistinguishable from P. lewisii in flower number, bud exposure, or any other character. The elaborated inflorescences of some plants in this part of California are striking; for now P. californicus and relatives are treated as part of a broadly defined P. lewisii. In her phylogenetic analysis based on ITS sequence, A. E. Weakley (2002) also found that P. californicus was not distinguished from P. lewisii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 478. FNA vol. 12, p. 483.
Parent taxa Hydrangeaceae > Philadelphus Hydrangeaceae > Philadelphus
Sibling taxa
P. coronarius, P. inodorus, P. lewisii, P. mearnsii, P. microphyllus, P. pubescens, P. serpyllifolius, P. texensis
P. coronarius, P. hirsutus, P. inodorus, P. mearnsii, P. microphyllus, P. pubescens, P. serpyllifolius, P. texensis
Synonyms P. hirsutus var. intermedius, P. hirsutus var. nanus, P. sharpianus, P. sharpianus var. parviflorus P. californicus, P. confusus, P. cordifolius, P. gordonianus, P. gordonianus var. columbianus, P. helleri, P. insignis, P. lewisii var. angustifolius, P. lewisii subsp. californicus, P. lewisii var. ellipticus, P. lewisii subsp. gordonianus, P. lewisii var. gordonianus, P. lewisii var. helleri, P. lewisii var. intermedius, P. lewisii var. oblongifolius, P. lewisii var. parvifolius, P. lewisii var. platyphyllus, P. oreganus, P. trichothecus, P. zelleri
Name authority Nuttall: Gen. N. Amer. Pl. 1: 301. (1818) Pursh: Fl. Amer. Sept. 1: 329. (1813)
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