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Pinos Altos Mountain bean

Santa Rita Mountain bean

Stems

erect, trailing, or twining, 5–50 cm.

stiff-branched, usually trailing, sometimes climbing, to 300 cm.

Leaves

stipules sometimes red-purple pigmented, appressed to stem, reflexed or distally spreading, ovate to ovate-lanceolate, 1.8–6.5(–8.5) × 0.7–3(–4) mm (larger distally);

petiole 1.8–3.4 cm;

rachis 0.4–1 cm;

stipels subulate to ovate or lanceolate, 1–2.5 mm;

leaflet blades usually linear to lanceolate, sometimes narrowly trullate, sometimes terminal leaflets with 2 basal lobes each side, lateral leaflets often ± distinctly round lobed basally on 1 or both sides, laterals 2–4.7 × 0.5–1.3(–2.8) cm, terminal 2.4–5.6 × 0.3–1.2(–2.7) cm, thin, base attenuate, apex round or acute, obscurely apiculate, surfaces sparsely pubescent abaxially, scabrous adaxially.

often unifoliolate on 1–5 nodes of new growth;

stipules spreading or reflexed, triangular to lanceolate, 2–4.5 × 0.5–2 mm, strigillose;

petiole reduced, shorter than terminal leaflet (often only pulvinus), 0.4–3.5 cm;

rachis 0.5–2 cm;

stipels oblong to ovate or triangular, 1.5–2.5 mm;

leaflet blades often variegated on midvein, usually ovate to widely ovate or trullate to widely trullate, sometimes lanceolate, laterals 2.5–7(–8.6) × 1–5(–6.5) cm, terminal 3–8(–10.5) × 0.1–0.6(–1) cm, leathery, base oblique, cuneate or truncate, apex acute, strongly apiculate, surfaces glabrous or sparsely covered with incumbent and uncinate hairs along veins and margins.

Inflorescences

2–18 cm;

axis sparsely covered with minute, uncinate hairs;

rachis 0.2–0.6 cm, with 1–3 (usually 1-flowered) nodes;

bracts lanceolate, 4 × 1.5 mm, 4–6-veined.

to 85 cm;

main axis glabrous or sparsely covered with uncinate hairs, often also with incumbent hairs;

rachis to 45 cm, with 6–25 biflorous nodes or with additional flowering axes (sometimes proximal nodes subopposite);

primary bracts triangular, 1–2.5(–3.5) × 0.5–1 mm, 3-veined.

Peduncles

2–12 cm.

often with lateral inflorescences, to 40 cm.

Pedicels

to 10 mm, shorter than flowers, uncinate-pubescent;

bracteoles persistent, ovate or obsolete, 0.1–0.5 mm.

3–10 mm, glabrous or sparsely to densely covered with uncinate hairs;

bracteoles often persistent, triangular to lanceolate, 0.5–1 mm, apex ciliate.

Flowers

calyx campanulate-tubular, 5–8 mm, adaxial proximal portion rounded, papillate, covered with minute, uncinate hairs;

abaxial lobes lanceolate;

lateral lobes usually triangular, sometimes falcate;

adaxial lobes connate nearly throughout;

corolla violet, lavender-pink, or purple, 15–25 mm;

banner obovate or spatulate, 12–16 mm, apex obtuse, emarginate;

wings obovate, 15–27 mm, rounded-auriculate on distal margin at base;

keel 10 mm;

ovary linear, 6–7.5 mm, glabrous, marginally scabrous.

calyx campanulate, 3–4 mm, glabrous or sparsely covered with incumbent and uncinate hairs;

abaxial lobes triangular;

lateral lobes triangular;

adaxial lobes ± connate;

corolla lavender to purple or violet, 13 mm;

banner obovate to oblong, 10 mm, apex emarginate, outer surface often sparsely to densely covered with appressed hairs;

wings obovate to oblong, 10–13 mm;

keel 6 mm;

ovary linear, 4–5 mm, glabrous.

Legumes

horizontal to pendent, terete, linear-straight to slightly falcate, 35–45 × 3–4 mm, elastically dehiscent, valves leathery, glabrous.

pendent, compressed, oblong-falcate, (25–)35–53 × 6–11 mm, elastically dehiscent, valves leathery, glabrous, stipe to 5.5 mm.

Seeds

8–11, usually brown to light brown or grayish green, punctate and mottled black, sometimes light green, oblong or reniform, 2–3.7 × 1.7–2.6 mm, smooth;

hilum ovate-oblong, 0.4–0.5 mm.

3–6, grayish or light brown, mottled and streaked with black, oblong to transversely oblong, 4–7.5 × 4–5.7 mm;

hilum ovate to lanceolate, 1.2 mm, smooth.

Vines

perennial, with deep-seated, oblong or spherical tuberous taproots.

perennial, with large, tuberous taproots.

2n

= 22.

= 22.

Phaseolus parvulus

Phaseolus ritensis

Phenology Flowering Jul–Sep. Flowering Jul–Oct.
Habitat Pinus-Pseudotsuga or pine-oak forests, wet meadows. Rocky volcanic slopes or ash, sandy or gravelly soils, pine-oak forests.
Elevation 1500–3000 m. (4900–9800 ft.) 1300–2700 m. (4300–8900 ft.)
Distribution
from FNA
AZ; NM; Mexico (Chihuahua, Durango, Jalisco, Nayarit, Sinaloa, Sonora, Zacatecas)
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; TX; Mexico (Chihuahua, Durango, Guanajuato, Jalisco, Nayarit, Sinaloa, Sonora, Zacatecas)
[BONAP county map]
Discussion

Phaseolus parvulus is known from Apache, Cochise, Coconino, Graham, Pima, and Santa Cruz counties in Arizona, and Catron, Grant, Hidalgo, and Sierra counties in New Mexico.

Phaseolus parvulus is a relatively small and unique wild bean characterized by its small, radishlike root, erect to trailing habit, and tubular calyx.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Phaseolus ritensis is known from Apache, Cochise, Gila, Graham, Maricopa, Pima, and Santa Cruz counties in Arizona, Catron County in New Mexico, and El Paso and Jeff Davis counties in Texas.

Roots of Phaseolus ritensis are reportedly used as a base for fermenting, as a purgative, and as a stimulant. The foliage has been used for animal forage (G. P. Nabhan et al. 1980).

Phaseolus ritensis and P. maculatus are similar in appearance, and both species, in flower, have been confused. In describing P. ritensis, M. E. Jones sensed the similarity with P. maculatus (there P. retusus) and explained that P. ritensis rarely has obtuse, never retuse, leaflets tips; bracts are minute, not conspicuous and persistent as in P. maculatus, and that the fruit of P. ritensis is distinctly falcate and long-stipitate. His diagnosis specified quite neatly the differences between the species. G. P. Nabhan et al. (1980) later revealed new distinguishable features between both taxa, such as the eophylls morphology, number of leaflets on the third node, leaf color, inflorescence length, fruit length, fruit valves and their distinct degrees of dehiscence, and seed dimensions.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Phaseolus Fabaceae > subfam. Faboideae > Phaseolus
Sibling taxa
P. acutifolius, P. angustissimus, P. filiformis, P. leptostachyus, P. maculatus, P. polystachios, P. ritensis, P. scabrellus, P. sinuatus, P. smilacifolius, P. texensis
P. acutifolius, P. angustissimus, P. filiformis, P. leptostachyus, P. maculatus, P. parvulus, P. polystachios, P. scabrellus, P. sinuatus, P. smilacifolius, P. texensis
Synonyms Alepidocalyx parvulus P. maculatus subsp. ritensis
Name authority Greene: Bot. Gaz. 6: 217. (1881) M. E. Jones: Contr. W. Bot. 12: 14. (1908)
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