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Pinos Altos Mountain bean

Metcalf bean, spotted bean

Stems

erect, trailing, or twining, 5–50 cm.

coarse, trailing or climbing, to 600 cm.

Leaves

stipules sometimes red-purple pigmented, appressed to stem, reflexed or distally spreading, ovate to ovate-lanceolate, 1.8–6.5(–8.5) × 0.7–3(–4) mm (larger distally);

petiole 1.8–3.4 cm;

rachis 0.4–1 cm;

stipels subulate to ovate or lanceolate, 1–2.5 mm;

leaflet blades usually linear to lanceolate, sometimes narrowly trullate, sometimes terminal leaflets with 2 basal lobes each side, lateral leaflets often ± distinctly round lobed basally on 1 or both sides, laterals 2–4.7 × 0.5–1.3(–2.8) cm, terminal 2.4–5.6 × 0.3–1.2(–2.7) cm, thin, base attenuate, apex round or acute, obscurely apiculate, surfaces sparsely pubescent abaxially, scabrous adaxially.

stipules spreading, later reflexed, ovate to lanceolate, often slightly lobed basally, 5–10 × 1–2.5 mm;

petiole 1.5–6.5 cm;

rachis 1.4–3.6 cm;

stipels ovate to triangular, 1.3–3.5 mm, 3-veined;

leaflet blades often variegated on midvein, widely ovate to very widely depressed-ovate or rhombic, laterals 3–7 × 2.5–5 cm, terminal 3–9.5 × 3–7.4 cm, leathery, base rounded to subcordate or subtruncate to cuneate, apex obtuse to rounded, rarely acute, retuse to emarginate, strongly apiculate, surfaces covered with minute uncinate hairs, margins and often abaxial surfaces, along prominent veins with incumbent hairs.

Inflorescences

2–18 cm;

axis sparsely covered with minute, uncinate hairs;

rachis 0.2–0.6 cm, with 1–3 (usually 1-flowered) nodes;

bracts lanceolate, 4 × 1.5 mm, 4–6-veined.

to 50 cm;

main axis sparsely to densely covered with minute uncinate hairs;

rachis 5–24 cm, with 6–20 nodes, often proximal nodes subopposite, biflorous;

primary bracts usually lanceolate, sometimes orbiculate, (2–)3–6.5(–8) × 0.5–1.5(–2) mm, 3–6-veined, glabrous or covered by uncinate and appressed hairs.

Peduncles

2–12 cm.

often with lateral inflorescences, sometimes with a floral node close to base, to 28 cm.

Pedicels

to 10 mm, shorter than flowers, uncinate-pubescent;

bracteoles persistent, ovate or obsolete, 0.1–0.5 mm.

3.5–7(–10) mm, covered with uncinate hairs, often with scattered incumbent hairs;

bracteoles often persistent, lanceolate to oblanceolate, often clavate, 0.8–2 mm.

Flowers

calyx campanulate-tubular, 5–8 mm, adaxial proximal portion rounded, papillate, covered with minute, uncinate hairs;

abaxial lobes lanceolate;

lateral lobes usually triangular, sometimes falcate;

adaxial lobes connate nearly throughout;

corolla violet, lavender-pink, or purple, 15–25 mm;

banner obovate or spatulate, 12–16 mm, apex obtuse, emarginate;

wings obovate, 15–27 mm, rounded-auriculate on distal margin at base;

keel 10 mm;

ovary linear, 6–7.5 mm, glabrous, marginally scabrous.

calyx campanulate, 4–5.5 mm, sparsely strigillose to short-strigose;

abaxial lobes narrowly triangular;

lateral lobes triangular;

adaxial lobes connate, often partially divided;

corolla reddish, violet, or bright pink to purple, 10–15 mm;

banner oblong to obovate, 10 mm, apex emarginate, adaxial surface often covered with appressed hairs;

wings obovate to widely obovate, 10–15 mm;

keel 7 mm;

ovary oblong, 4–5 mm, glabrous.

Legumes

horizontal to pendent, terete, linear-straight to slightly falcate, 35–45 × 3–4 mm, elastically dehiscent, valves leathery, glabrous.

pendent, compressed, oblong, somewhat curved, or obovate (when 1-seeded), 30–74 × 12–17 mm, sometimes with constriction in proximal 1/2, weakly dehiscent or sometimes elastically dehiscent, valves leathery, glabrous, sessile.

Seeds

8–11, usually brown to light brown or grayish green, punctate and mottled black, sometimes light green, oblong or reniform, 2–3.7 × 1.7–2.6 mm, smooth;

hilum ovate-oblong, 0.4–0.5 mm.

1–5, brown, mottled and often streaked with black, suborbicular to subquadrate, (6–)8–13 mm diam., smooth;

hilum ovate to lanceolate or oblong, 1.3 mm.

Vines

perennial, with deep-seated, oblong or spherical tuberous taproots.

perennial, with large, tuberous, woody taproots.

2n

= 22.

= 22.

Phaseolus parvulus

Phaseolus maculatus

Phenology Flowering Jul–Sep. Flowering Jun–Oct.
Habitat Pinus-Pseudotsuga or pine-oak forests, wet meadows. Rocky slopes of deep canyons, valley bottoms, along streams, well-drained, sandy soils, open pine-oak or oak forests, grasslands, dry scrub.
Elevation 1500–3000 m. (4900–9800 ft.) 1400–2700 m. (4600–8900 ft.)
Distribution
from FNA
AZ; NM; Mexico (Chihuahua, Durango, Jalisco, Nayarit, Sinaloa, Sonora, Zacatecas)
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; TX; Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Phaseolus parvulus is known from Apache, Cochise, Coconino, Graham, Pima, and Santa Cruz counties in Arizona, and Catron, Grant, Hidalgo, and Sierra counties in New Mexico.

Phaseolus parvulus is a relatively small and unique wild bean characterized by its small, radishlike root, erect to trailing habit, and tubular calyx.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Phaseolus maculatus is found throughout Arizona, except the western tier of counties, in the southwestern quarter of New Mexico, and in Brewster, Jeff Davis, Presidio, and Terrell counties in Texas.

Phaseolus maculatus is characterized by leaves with rigid margins and relatively large, passively or weakly dehiscent pods.

Phaseolus maculatus is closely related to P. ritensis. Both possess deep and enlarged taproots, trailing habits, leathery leaves with short petioles, and long inflorescences with few to many flowers. It has also been reported as P. ritensis for its useful roots, foliage (forage), and seed (G. P. Nabhan et al. 1980).

Phaseolus retusus Bentham (1839) is an illegitimate name that pertains here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Phaseolus Fabaceae > subfam. Faboideae > Phaseolus
Sibling taxa
P. acutifolius, P. angustissimus, P. filiformis, P. leptostachyus, P. maculatus, P. polystachios, P. ritensis, P. scabrellus, P. sinuatus, P. smilacifolius, P. texensis
P. acutifolius, P. angustissimus, P. filiformis, P. leptostachyus, P. parvulus, P. polystachios, P. ritensis, P. scabrellus, P. sinuatus, P. smilacifolius, P. texensis
Synonyms Alepidocalyx parvulus P. metcalfei
Name authority Greene: Bot. Gaz. 6: 217. (1881) Scheele: Linnaea 21: 465. (1848)
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