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upsidedown grass

Habit Plants cespitose. Plants perennial, rhizomatous; monoecious.
Culms

25-95 cm, generally decumbent and rooting at the nodes.

annual, not woody, solid or hollow.

Sheaths

glabrous, extensively overlapping;

ligules 1-2 mm;

pseudopetioles 8-60 mm;

blades 7-30 cm long, 2-6.5 cm wide, narrowly elliptic to obovate, often acuminate, lacking intercostal fibrous bands, sometimes whitened beneath, lateral veins diverging from the midvein at a 4-8° angle.

Leaves

mostly cauline, evidently distichous;

sheaths open;

auricles absent;

abaxial ligules absent;

adaxial ligules membranous, ciliate;

pseudopetioles prominent, twisted so the blades are resupinate;

blades with the lateral veins evident, straight, diverging obliquely from the central vein;

mesophyll nonradiate;

adaxial palisade layer absent;

fusoid cells well developed, large;

arm cells weakly to moderately developed;

Kranz anatomy not developed;

midribs complex; inflated interstomatal cells present;

bulliform cells poorly developed or absent;

stomates with parallel-sided to dome-shaped subsidiary cells;

bicellular microhairs absent;

papillae absent.

Panicles

10-40 cm, sparsely flowered;

branches solitary, with uncinate hairs, usually tipped with a staminate spikelet.

Inflorescences

panicles, rachises and branches sometimes disarticulating, hairy, hairs uncinate.

Spikelets

unisexual, with 1 floret, mostly in staminate-pistillate pairs on short branchlets, some pistillate spikelets solitary.

Caryopses

hila subequal to the caryopses, linear;

endosperm hard, without lipid;

embryos small;

epiblasts present;

scutellar cleft present but short;

mesocotyl internode absent;

embryonic leaf margins overlapping, x = 12.

Staminate

spikelets 2.5-3.5 mm, on 4-11 mm pedicels, subtending the pistillate spikelets, purple;

lower glumes 1-2 mm;

upper glumes 1.5-3.2 mm, 1- or 3-veined;

lemmas 2.5-3.5 mm;

paleas about 3/4 the length of the lemmas;

anthers 0.9-1.1 mm.

spikelets smaller than the pistillate spikelets, short- to long-pedicellate;

glumes 1-2, membranous, shorter than the floret;

lodicules absent or 3 and minute, elliptic, glabrous, veinless;

anthers 6.

Pistillate

spikelets 7.5-12 mm, diverging slightly from the branches;

glumes brown;

lower glumes 5-7 mm, 5-7-veined;

upper glumes 6-8 mm, 3-5-veined;

lemmas 7.5-12 mm, linear-oblong, abruptly short-beaked, with uncinate hairs nearly to the base;

paleas equaling the lemmas.

spikelets sessile or shortly pedicellate;

glumes 2, shorter than the floret;

lemmas tubular or inflated, covered wholly or in part with uncinate hairs, unawned;

paleas well developed;

lodicules absent;

ovaries glabrous, without an apical appendage;

styles 1, with 3 branches.

2n

= 24.

Pharus glaber

Poaceae subfam. pharoideae

Distribution
from USDA
Discussion

Pharus glaber grows on limestone-influenced sand in the hammocks of central Florida. Only two remaining populations are known in the United States, but the species is still widely present elsewhere in the Neotropics. Hitchcock (1951) erroneously referred this species to Pharus parvifolius Nash, which differs primarily in the presence of intercostal fibrous bands on the adaxial surfaces of the leaf blades.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The Pharoideae include one tribe, the Phareae, three genera, and twelve species. It is pantropical, but in the Americas, it is represented by one genus, Pharus, that extends from Florida to Uruguay and Argentina. The Pharoideae is a basal lineage of the Poaceae, and the first subfamily in which an adaxial ligule and true spikelets are found.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 24, p. 13. FNA vol. 24, p. 11. Author: Grass Phylogeny Working Group;.
Parent taxa Poaceae > subfam. Pharoideae > tribe Phareae > Pharus Poaceae
Subordinate taxa
Name authority Kunth L.G. Clark & Judz.
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