Phalaris lemmonii |
Poaceae subfam. pooideae |
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Lemmon's Canary grass |
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Habit | Plants annual. | Plants annual or perennial; sometimes matlike, sometimes cespitose, sometimes stoloniferous, sometimes rhizomatous. | ||||||||||||||||||||||||||||||||||||
Culms | (7)25-150 cm. |
usually hollow, sometimes solid. |
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Leaves | distichous; sheaths usually open to the base, varying to closed for nearly their full length; auricles present or absent; abaxial ligules absent; adaxial ligules scarious or membranous, sometimes puberulent or scabridulous, usually not ciliate, cilia sometimes shorter than the base; pseudopetioles rarely present; blades usually linear, sometimes broadly so, venation parallel; cross sections non-Kranz, mesophyll nonradiate, adaxial palisade layer absent, fusoid and arm cells usually absent; midribs usually simple; adaxial bulliform cells present; stomates with parallel-sided subsidiary cells; epidermes usually lacking bicellular microhairs, sometimes with unicellular microhairs, papillae usually absent, when present, rarely more than 1 per cell. |
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Panicles | (2)3-20 cm long, 0.6-1.5 cm wide, cylindrical, evidently branched below; branches to 2 cm, spikelets borne singly, not clustered. |
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Inflorescences | usually terminal, panicles, spikes, or racemes, usually ebracteate; disarticulation usually below the florets, sometimes below the glumes, at the rachis nodes, or at the inflorescence bases. |
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Spikelets | homogamous, with (2)3 florets, terminal floret bisexual; disarticulation above the glumes, beneath the sterile florets. |
usually bisexual, infrequently unisexual or mixed, usually laterally compressed or not compressed, occasionally dorsally compressed, with 1-30 sexual florets, distal floret(s) often reduced, infrequently spikelets with 1-2 reduced or staminate basal florets and a single terminal sexual floret. |
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Glumes | 4.5-6.7 mm long, 0.9-1.1 mm wide, acuminate, keels not or only slightly winged, wings to 0.2 mm wide, keels, lateral veins, and adjacent surfaces scabrous; sterile florets (1)2, 1-1.6 mm, densely appressed-hairy; bisexual florets 2.7-5.1 mm long, 1.2-1.6 mm wide, shiny, stramineous to gray-brown, mostly hairy with spreading hairs, apices glabrous, strongly acuminate to beaked; anthers 0.7-2 mm. |
usually 2, upper or lower glumes sometimes absent, rarely both glumes absent; lemmas without uncinate hairs, awned or not, awns single, basal to apical; paleas usually well-developed, sometimes reduced or absent; lodicules 2(3), usually lanceolate and broadly membranous distally, rarely truncate and fleshy, usually not veined or obscurely veined, sometimes distinctly veined, sometimes ciliate; anthers (1, 2)3; ovaries glabrous or sometimes hairy distally, sometimes with an apical appendage; haustorial synergids absent; styles (1)2 (-4), bases close together, sometimes fused. |
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Caryopses | hila linear, elliptic, ovate, or punctate; endosperm usually hard, sometimes soft or liquid, with or without lipids, starch grains compound or simple; embryos less than 1/2 the length of the caryopses; epiblasts usually present; scutellar cleft usually absent; mesocotyl internode usually absent; embryonic leaf margins overlapping, x = 7, 10. |
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Ligules | 1.5-8 mm, acute; blades to 14 cm long, 1-8 mm wide, smooth, shiny, sometimes revolute. |
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2n | = 14. |
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Phalaris lemmonii |
Poaceae subfam. pooideae |
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Distribution |
CA
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Discussion | Phalaris lemmonii is native to California, but it has also been found in Victoria, Australia. It grows in moist areas, and appears to hybridize with both P. caroliniana and P. angusta (Baldini 1995). The strongly beaked tips of the bisexual florets are a useful distinguishing feature. Beecher Crampton noted on one unusually small specimen (UTC 230918) that it was the vernal pool ecotype of the species. He did not publish his observations. Anderson (1961) and Baldini (1995) distinguished Phalaris lemmonii from P. platensis Henrard ex Wacht., a narrowly distributed South American taxon, arguing that it was slightly longer in the length of its ligules, glumes, florets, and anthers, but many California specimens fall within the range given for P. platensis rather than that for P. lemmonii. Phalaris lemmonii is the older name so, if further research shows that the two species should be combined, P. lemmonii will remain as the correct name for plants from the Flora region. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The subfamily Pooideae includes approximately 3300 species, making it the largest subfamily in the Poaceae. It reaches its greatest diversity in cool temperate and boreal regions, extending across the tropics only in high mountains. The circumscription and relationships of tribes within the Pooideae are unsettled (see, for example, Catalan et al. 1997, 2004; Soreng and Davis 1998). In this flora, some previously recognized tribes have been combined with the Poeae. Recognition of some of these as subtribes is well supported; among these is the Hainardieae Greuter (which, at the subtribal level, is called the Parapholiinae Caro). Members of other traditional tribal groupings, such as the Aveneae Dumort., appear to be widely dispersed within the Poeae sensu lato. Further work will probably support the division of the expanded Poeae into additional tribes; there is as yet no clear indication as to what the boundaries of such tribes should be. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24. | FNA vol. 24, p. 57. | ||||||||||||||||||||||||||||||||||||
Parent taxa | Poaceae > subfam. Pooideae > tribe Poeae > Phalaris | Poaceae | ||||||||||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||||||||||
Name authority | Vasey | Benth. | ||||||||||||||||||||||||||||||||||||
Web links |