r enouée robuste, stout dotted smartweed, stout smartweed
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Plants perennial, 3–20 dm; roots also sometimes arising from proximal nodes; rhizomes present, stolons sometimes present. |
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ascending, usually branched proximally, scarcely ribbed, glabrous, glandular-punctate; branches sometimes creeping and rooting at nodes. |
ascending to erect or, rarely, prostrate, usually branched, sometimes simple, unarmed. |
ocrea light brown, cylindric, 10–15 mm, chartaceous, base inflated, margins truncate, ciliate with bristles 3–12 mm, surface strigose, glandular-punctate; petiole 0.2–2 cm, glandular-punctate; blade without dark triangular or lunate blotch adaxially, lanceolate to elliptic-lanceolate, 4–20 × 2–4.5 cm, base tapered, margins antrorsely strigose, apex acute to acuminate, faces glabrous or main veins scabrous, glandular-punctate. |
ocrea chartaceous, rarely foliaceous distally, margins eciliate or ciliate; petiole not winged, not auriculate; blade sometimes with dark triangular or lunate blotch adaxially, lanceolate to narrowly elliptic, base tapered, acute, rounded, or cordate, margins entire. |
terminal and axillary, erect, uninterrupted, 20–80 × 5–10 mm; peduncle 5–40 mm, glandular-punctate; ocreolae usually overlapping, margins eciliate or proximal ones sometimes ciliate with bristles to 1 mm. |
terminal or terminal and axillary, spikelike, uninterrupted or interrupted. |
ascending to spreading, 2–5 mm. |
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2–4 per ocreate fascicle, homostylous; perianth greenish proximally, white distally, glandular-punctate with punctae ± uniformly distributed, slightly accrescent; tepals 5, connate ca.1/3 their length, obovate, 3.2–4.2 mm, veins prominent or not, not anchor-shaped, margins entire, apex obtuse to rounded; stamens 6–8, included; anthers pink or red, elliptic; styles 3, connate proximally. |
homostylous or heterostylous, articulation swollen or not; perianth campanulate; tepals 4–5, connate 1/4–2/3 their length; stamens 5–8; styles deciduous, 2–3, included or exserted (exserted syles and stamens in heterostylous species), spreading. |
included or apex exserted, dark brown to brownish black, 3-gonous, 2.7–3.6 × 2–2.5 mm, shiny, smooth. |
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Flowering Jul–Oct. |
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Peaty shores, often in water on coastal plain or near coast |
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0-1500 m (0-4900 ft) |
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CT; DE; FL; IN; KY; MA; MD; ME; MI; MO; NH; NJ; NY; OH; PA; RI; TX; VA; NS; ON; QC; Central America; South America; West Indies
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Nearly worldwide; especially n temperate regions |
Persicaria robustior often is synonymized with P. punctata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 60 (16 in the flora). Members of sect. Persicaria frequently grow in moist or inundated habitats. Many have evolved responses to submergence that allow them to survive extended periods of inundation (R. S. Mitchell 1976). Hybridization often is implicated for the taxonomic difficulties in the section. However, J. Timson (1965) concluded that, at least among annual European species, hybridization is rare due to autogamous breeding systems. Mitchell (1971) demonstrated the usefulness of leaf morphology in distinguishing major groups among the native North American smartweeds. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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1. Some or all ocreae foliaceous and green distally | → 2 |
1. All ocreae chartaceous and hyaline, tan, brown, or reddish brown throughout, never foliaceous and green distally | → 3 |
2. Plants annual; rhizomes and stolons absent; leaf blades ovate, 3-17 cm wide | P. orientalis |
2. Plants perennial; rhizomes or stolons usually present; leaf blades ovate-lanceolate to elliptic or oblong-lanceolate, 1-6(-8) cm wide | P. amphibia |
3. Perianths glandular-punctate | → 4 |
3. Perianths not glandular-punctate | → 9 |
4. Achenes minutely roughened, dull; axillary inflorescences sometimes enclosed in ocreae | P. hydropiper |
4. Achenes smooth, shiny; inflorescences never enclosed in ocreae | → 5 |
5. Outer tepals with anchor-shaped veins; achenes discoid | P. lapathifolia |
5. Outer tepals without anchor-shaped veins; achenes 3-gonous or biconvex | → 6 |
6. Ocreae margins eciliate; achenes biconvex; styles 2 | P. glabra |
6. Ocreae margins ciliate with bristles 2-12 mm; achenes usually 3-gonous, rarely biconvex; styles 2-3 | → 7 |
7. Punctae confined to bases of perianths and sometimes on inner tepals | P. hydropiperoides |
7. Punctae ± uniformly distributed over perianths | → 8 |
8. Inflorescences interrupted; ocreolae mostly not overlapping, margins mostly ciliate with hairs to 2 mm; leaf blades 0.6-2.4 cm wide | P. punctata |
8. Inflorescences uninterrupted; ocreolae usually overlapping, margins mostly eciliate or proximal sometimes with hairs to 1 mm; leaf blades 2-4.5 cm wide | P. robustior |
9. Peduncles stipitate-glandular | → 10 |
9. Peduncles not stipitate-glandular | → 14 |
10. Plants perennial; rhizomes or stolons usually present; inflorescences terminal | P. amphibia |
10. Plants annual; rhizomes and stolons absent; inflorescences terminal and axillary | → 11 |
11. Stems hirsute proximally; margins of ocreae ciliate with bristles 2-7 mm | P. careyi |
11. Stems glabrous proximally; margins of ocreae eciliate or ciliate with bristles less than 1 mm | → 12 |
12. Outer tepals with anchor-shaped veins; tepals 4(-5); inflorescences mostly arching or nodding | P. lapathifolia |
12. Outer tepals without anchor-shaped veins; tepals 5; inflorescences erect or, rarely, nodding | → 13 |
13. Flowers homostylous; achenes without central hump on 1 side | P. pensylvanica |
13. Flowers heterostylous; achenes usually with central hump on 1 side | P. bicornis |
14. Plants perennial; rhizomes or stolons usually present | → 15 |
14. Plants annual; rhizomes and stolons absent | → 19 |
15. Achenes biconvex; styles 2 | → 16 |
15. Achenes 3-gonous; styles 3 | → 17 |
16. Perianth greenish white or white to pink; surfaces of ocreae glabrous, usually obscurely glandular-punctate | P. glabra |
16. Perianth roseate to red; surfaces of ocreae glabrous or appressed-pubescent to hirsute, not glandular-punctate | P. amphibia |
17. Proximal part of ocreae strigose, or ocreae glabrous | P. hydropiperoides |
17. Proximal part of ocreae hirsute, or strigose and with loosely ascending to spreading hairs at least proximally | → 18 |
18. Bases of leaf blades rounded to cordate; stems brownish-hirsute on internodes | P. hirsuta |
18. Bases of leaf blades tapered to truncate; stems glabrous or loosely appressed- to spreading-hirsute near nodes | P. setacea |
19. Margins of ocreae without bristles or with bristles to 1 mm; ocreolae mostly overlapping; achenes discoid, rarely 3-gonous | → 20 |
19. Margins of ocreae ciliate with bristles (0.2-)1-12 mm, if bristles less than 1 mm then ocreolae not overlapping; achenes discoid, biconvex, or 3-gonous | → 22 |
20. Outer tepals with anchor-shaped veins; tepals 4(-5); inflorescences mostly arching or nodding | P. lapathifolia |
20. Outer tepals without anchor-shaped veins; tepals 5; inflorescences mostly erect, rarely nodding | → 21 |
21. Flowers homostylous; achenes without central hump on 1 side | P. pensylvanica |
21. Flowers heterostylous; achenes usually with central hump on 1 side | P. bicornis |
22. Inflorescences not dense; ocreolae not overlapping proximally, usually overlapping distally; leaf blades linear to linear-lanceolate | P. minor |
22. Inflorescences dense; ocreolae mostly overlapping; leaf blades narrowly ovate or ovate-lanceolate to linear-lanceolate | → 23 |
23. Bristles of ocreolae 0.2-1.3(-2) mm; achenes discoid, biconvex, or 3-gonous; styles 2-3 | P. maculosa |
23. Bristles of ocreolae (0.5-)1-4(-6) mm; achenes 3-gonous; styles 3 | P. longiseta |
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FNA vol. 5, p. 584. |
FNA vol. 5, p. 581. |
Polygonaceae > subfam. Polygonoideae > Persicaria > sect. Persicaria |
Polygonaceae > subfam. Polygonoideae > Persicaria |
P. amphibia, P. arifolia, P. bicornis, P. bungeana, P. capitata, P. careyi, P. chinensis, P. glabra, P. hirsuta, P. hydropiper, P. hydropiperoides, P. lapathifolia, P. longiseta, P. maculosa, P. meisneriana, P. minor, P. nepalensis, P. orientalis, P. pensylvanica, P. perfoliata, P. punctata, P. sagittata, P. setacea, P. virginiana, P. wallichii |
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P. amphibia, P. bicornis, P. careyi, P. glabra, P. hirsuta, P. hydropiper, P. hydropiperoides, P. lapathifolia, P. longiseta, P. maculosa, P. minor, P. orientalis, P. pensylvanica, P. punctata, P. robustior, P. setacea |
Polygonum punctatum var. robustius, P. punctata var. robustior, Polygonum punctatum var. majus, Polygonum robustius |
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(Small) E. P. Bicknell: Bull. Torrey Bot. Club 36: 455. (1909) |
unknown |
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