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Bolander's Yampah, olasi

Howell's Yampah

Habit Herbs 15–90 cm; roots tuberous, 1 or 2–3-clustered, 1–7 cm. Herbs 80–160 cm; roots fibrous, ± thickened, 15+-clustered, 4–15 cm.
Leaves

basal 2–4 pinnate;

petiole 2–15 cm;

blade deltate, ovate, or ovate-lanceolate, 10–20 cm, most primary leaflets dissected, ultimate divisions monomorphic or dimorphic (terminal more elongate than laterals), oblong to filiform, 5–60 mm (terminal sometimes to 80 mm), margins usually toothed or lobed;

cauline leaves usually 1-ternate or 1-ternate-pinnate.

basal 2-pinnate;

petiole 10–20 cm;

blade ± lanceolate, 30–50 cm, most primary leaflets dissected, ultimate divisions monomorphic, widely lanceolate to ovate, 20–70 × 10–40 mm, margins deeply toothed or lobed;

cauline leaves (1–)2-pinnate.

Peduncles

2–20 cm.

5–25 cm.

Umbels

involucral bracts 8–12, linear to narrowly lanceolate;

rays 9–23, 1–2 cm, subequal, ascending or spreading-ascending;

involucel bractlets 4–10, obovate or lanceolate to linear-lanceolate, 3–9 mm, widely scarious-margined or entirely scarious, apex acuminate;

umbellets convex, 18–30-flowered.

involucral bracts 8–12, linear to linear-spatulate;

rays (15–)17–20(–32), 2–7 cm, ± equal, spreading-ascending;

involucel bractlets ± 10, lanceolate, 5–12 mm;

umbellets convex, 25–30-flowered.

Pedicels

2–5 mm.

5–15 mm.

Flowers

petals 1-veined;

styles 2 mm, spreading and usually recurved.

petals 5–7-veined;

styles less than 0.5 mm, ± erect.

Schizocarps

oblong;

mericarps 4–6 mm;

ribs threadlike;

oil ducts 2–3 in intervals, 4 on commissure.

oblong;

mericarps 4–6 mm;

ribs conspicuously corky;

oil ducts 1 in intervals, 2 on commissure.

2n

= 40.

Perideridia bolanderi

Perideridia howellii

Phenology Flowering Jul–Sep; fruiting Sep–Oct.
Habitat Moist meadows, ravines, stream banks.
Elevation 100–1600 m. [300–5200 ft.]
Distribution
map from USDA
w United States
[WildflowerSearch map]
[BONAP county map]
map from FNA
CA; OR
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Perideridia bolanderi is distinguishable from all other species in the genus by the following combination of characters: ultimate divisions of the basal leaves dimorphic, or if homomorphic, less than 1 mm wide; bracts and bractlets broad, conspicuously scarious, and cuspidate or acuminate, or if only broadly scarious-margined and acuminate, the bracts persistent at maturity; and mericarps oblong with 2–3 oil ducts in the intervals. Although the two subspecies share this combination of characters, phylogenetic analysis of DNA sequence data shows that they do not form a monophyletic group, though this separation is only weakly supported (S. R. Downie et al. 2004).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Perideridia howellii is the most distinctive species in the genus and is easily separated from all other species of Perideridia by the combination of broad leaflets, conic stylopodium, and prominently corky fruit ribs. It is found in the Cascade Range and Sierra Nevada from Klamath County, Oregon, to Mariposa County, California, and in the Inner Coast and Klamath ranges from Douglas County, Oregon, to Sonoma County, California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Parent taxa Apiaceae > Perideridia Apiaceae > Perideridia
Sibling taxa
P. americana, P. bacigalupii, P. californica, P. erythrorhiza, P. gairdneri, P. howellii, P. kelloggii, P. lemmonii, P. leptocarpa, P. montana, P. oregana, P. parishii, P. pringlei
P. americana, P. bacigalupii, P. bolanderi, P. californica, P. erythrorhiza, P. gairdneri, P. kelloggii, P. lemmonii, P. leptocarpa, P. montana, P. oregana, P. parishii, P. pringlei
Subordinate taxa
P. bolanderi subsp. bolanderi, P. bolanderi subsp. involucrata
Key
1. Plants not glaucous; ultimate divisions of basal leaf blades 1–4 mm wide, dimorphic (terminal more elongate than laterals); involucral bracts and involucel bractlets entirely scarious, deciduous.
subsp. bolanderi
1. Plants glaucous; ultimate divisions of basal leaf blades 0.3–0.7 mm wide, monomorphic; involucral bracts and involucel bractlets widely scarious-margined, persistent.
subsp. involucrata
Synonyms Podosciadium bolanderi Carum howellii
Name authority (A. Gray) A. Nelson & J. F. Macbride: Bot. Gaz. 61: 33. (1916) (J. M. Coulter & Rose) Mathias: Brittonia 2: 244. (1936)
Source FNA vol. 13. Treatment authors: Fengjie Sun, Stephen R. Downie. FNA vol. 13. Treatment authors: Fengjie Sun, Stephen R. Downie.
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