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ditch-stonecrop, Virginia penthorum

ditch, penthorum family, stonecrop family

Stems

frequently pink or reddish, especially with age, 1–6(–8) dm, stipitate glands hyaline or reddish brown and often black- or purple-tipped.

simple or branched.

Leaves

petiole absent or 1–10 mm;

blade 2–18(–24) × 0.5–4(–5.5) cm, glabrous or sparsely stipitate-glandular abaxially and adaxially, hairs black- or purple-tipped.

cauline, alternate, simple;

stipules absent;

petiole present or absent;

blade margins sharply serrate or doubly serrate.

Inflorescences

secund, 2–7-branched, each branch (6–)10–25(–30)-flowered.

terminal, scorpioid racemes.

Pedicels

0.5–3 mm.

Flowers

hypanthium 0.5–1.5 × 1.5–3.5 mm;

sepals persistent, erect or spreading, unequal, 0.8–2 × 0.4–1 mm, margins entire or serrulate with 1–4 gland-tipped teeth per side;

filaments 1–2 mm;

anthers 0.7–1 mm;

pistil 3–4 mm;

stigmas often purple with age.

bisexual;

perianth and androecium hypogynous or perigynous;

hypanthium present;

sepals 5(–8), connate proximally;

petals absent or 1–8, distinct;

nectary disc absent;

stamens 10, distinct, adnate to rim of hypanthium;

anthers dehiscent by longitudinal slits;

pistils 1, (4–)5(–8)-carpellate, connate basally and laterally, partially adnate to hypanthium;

ovary superior or partially inferior, (4–)5(–8)-locular proximally;

placentation marginal;

ovules anatropous, bitegmic, crassinucellate;

styles 1 per carpel, distinct;

stigmas 1 per carpel, terminal, capitate.

Fruits

capsular, dehiscence circumscissile proximal to free portion of styles.

Seeds

0.5–0.7 × 0.2–0.3 mm, tubercles reddish or pinkish.

300–400, tan, yellowish brown, or pinkish, ellipsoid to subfusiform, funicular end more tapered;

embryo straight;

endosperm scant, cellular.

Perennial

herbs.

2n

= 18.

Penthorum sedoides

Penthoraceae

Phenology Flowering Jul–Oct.
Habitat Wet soils, stream banks, fresh-water marshes, margins of beaver ponds, pools in floodplain forests, shores, ditches
Elevation 0-700 m (0-2300 ft)
Distribution
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NY; OH; OK; OR; PA; RI; SC; SD; TN; TX; VA; VT; WA; WI; WV; BC; MB; NB; ON; QC
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[BONAP county map]
North America; Europe (Russia); Asia
[BONAP county map]
Discussion

The seeds of Penthorum sedoides were used by the Meskwaki to make cough medicine, and the leaves were used by the Cherokee as a potherb (D. E. Moerman 1998). The species is introduced in southern British Columbia, Oregon, and Washington, where it grows in cranberry bogs.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genus 1, species 2 (1 in the flora).

The position of Penthorum within Rosales has been disputed extensively. A. Cronquist (1981) considered it to be transitional between Crassulaceae and Saxifragaceae. He included it in Saxifragaceae, stating that Penthorum was not distinct enough from Crassulaceae and Saxifragaceae to warrant being treated as a distinct family. Placement of the genus by others has depended on the morphological, anatomical, and embryological traits emphasized. Molecular studies suggest that the genus is sister to Haloragaceae (D. R. Morgan and D. E. Soltis 1993; D. E. Soltis and P. S. Soltis 1997). Recent authors often have placed it in the monogeneric Penthoraceae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 8, p. 231. FNA vol. 8, p. 230. Author: Craig C. Freeman.
Parent taxa Penthoraceae > Penthorum
Subordinate taxa
Name authority Linnaeus: Sp. Pl. 1: 432. (1753) Rydberg ex Britton
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