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dark beardtongue, Whipple's beardtongue, Whipple's penstemon

Sierra beardtongue, Sierra penstemon

Habit Herbs. Herbs.
Stems

ascending to erect, (8–)20–65(–100) cm, glabrous or ± puberulent proximally, ± puberulent or glandular-pubescent distally, not glaucous.

decumbent, ascending, or erect, 5–65 cm, glabrous or retrorsely hairy, sometimes also glandular-pubescent distally, not glaucous.

Leaves

basal and cauline, not leathery, glabrous;

basal and proximal cauline 40–90(–130) × (5–)10–30(–75) mm, blade ovate to lanceolate or elliptic, base tapered to cuneate, margins entire or ± repand, denticulate, or dentate, apex obtuse to acute;

cauline 2–5 pairs, short-petiolate or sessile, 25–60(–85) × 3–15(–25) mm, blade lanceolate to oblanceolate, base tapered to clasping or cordate-clasping, margins entire, sometimes ± repand to denticulate, apex obtuse to acute.

basal and cauline, not leathery, glabrous;

basal and proximal cauline 5–140 × 3–20 mm, blade spatulate to oblanceolate or elliptic, base tapered, margins entire, apex rounded to obtuse or acute;

cauline 2–5 pairs, short-petiolate or sessile, 10–60(–70) × 1–14 mm, blade oblong to oblanceolate or lanceolate, base tapered to clasping, margins entire, apex obtuse to acute.

Thyrses

interrupted or continuous, secund, (2–)5–35 cm, axis sparsely to densely glandular-pubescent, verticillasters 2–5(–7), cymes 2–4-flowered, 2 per node;

proximal bracts lanceolate, 11–85 × 1–18 mm, margins entire or ± repand proximally;

peduncles and pedicels ascending to erect, glandular-pubescent.

interrupted, cylindric, 1–12 cm, axis ± glandular-pubescent, verticillasters 1–6, cymes (1 or)2–6-flowered, 2 per node;

proximal bracts lanceolate, 4–35 × 1–6 mm, margins entire;

peduncles and pedicels erect, ± glandular-pubescent.

Flowers

calyx lobes lanceolate, 7–10 × 1–2.2 mm, glandular-pubescent;

corolla brownish purple to bluish violet, sometimes ochroleucous, lavender, or white, with alternating white or lavender and purple nectar guides in dark-colored forms, with lavender nectar guides in light-colored forms, ventricose to ventricose-ampliate, 20–27(–30) mm, glandular-pubescent externally, sparsely white-villous internally abaxially, tube 5–8 mm, throat abruptly inflated, 8–10 mm diam., 2-ridged abaxially;

stamens included or longer pair reaching orifice, pollen sacs opposite, explanate, 1–1.4 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth;

staminode 12–15 mm, reaching orifice or exserted, 0.6–1.1 mm diam., tip straight to slightly recurved, distal 1–3 mm sparsely to densely villous, hairs yellow, to 1 mm;

style 12–16 mm.

calyx lobes oblong to lanceolate, 4–6 × 0.7–1.2 mm, glandular-pubescent;

corolla lavender to blue, violet, or purple, essentially without nectar guides, tubular-funnelform, 10–16 mm, glandular-pubescent externally, moderately yellow-pilose to yellow-lanate internally abaxially, tube 5–7 mm, throat slightly inflated, 3–5 mm diam., prominently 2-ridged abaxially;

stamens: longer pair reaching orifice or slightly exserted, pollen sacs opposite, navicular, 0.6–0.8(–1) mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth;

staminode 6–9 mm, included, 0.3–0.6 mm diam., tip straight to recurved, glabrous or distal 1–3 mm sparsely to moderately pilose, hairs yellow, to 0.5 mm;

style 9–12 mm.

Capsules

6–9 × 4–5 mm, glandular-puberulent distally.

4–6 × 2.5–3.5 mm, glabrous.

2n

= 16.

Penstemon whippleanus

Penstemon heterodoxus

Phenology Flowering Jun–Aug(–Sep).
Habitat Rocky slopes in subalpine forests, alpine meadows.
Elevation 2100–3700 m. (6900–12100 ft.)
Distribution
from FNA
AZ; CO; ID; MT; NM; UT; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV
[WildflowerSearch map]
[BONAP county map]
Discussion

Penstemon whippleanus is known from forests and meadows of the Southern and Central Rocky mountains and westward into the Wasatch Mountains and Utah Plateaus. Plants with light-colored corollas often are found growing next to plants with dark-colored corollas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Penstemon heterodoxus comprises three partially sympatric varieties centered in the Sierra Nevada of eastern California.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Thyrse axes usually sparsely glandular-pubescent; n California.
var. shastensis
1. Thyrse axes moderately to densely glandular-pubescent; e California, wc Nevada.
→ 2
2. Stems 5–25 cm; basal and proximal cauline leaves 5–50 × 3–10 mm; verticillasters (1–)3; ec California, wc Nevada.
var. heterodoxus
2. Stems 15–35 cm; basal and proximal cauline leaves (17–)20–75 × 6–18 mm; verticillasters 2–6; se California.
var. cephalophorus
Source FNA vol. 17, p. 227. FNA vol. 17, p. 207.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. heterodoxus var. cephalophorus, P. heterodoxus var. heterodoxus, P. heterodoxus var. shastensis
Name authority A. Gray: Proc. Amer. Acad. Arts 6: 73. (1862) — (as Pentstemon) A. Gray: in A. Gray et al., Syn. Fl. N. Amer. 2(1): 269. (1878) — (as Pentstemon)
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