dark beardtongue, Whipple's beardtongue, Whipple's penstemon
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Alberta beardtongue, Alberta penstemon
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Herbs. |
Herbs. |
ascending to erect, (8–)20–65(–100) cm, glabrous or ± puberulent proximally, ± puberulent or glandular-pubescent distally, not glaucous. |
ascending to erect, (10–)15–38 cm, glabrous or sparsely retrorsely hairy, sometimes also sparsely glandular-pubescent. |
basal and cauline, not leathery, glabrous; basal and proximal cauline 40–90(–130) × (5–)10–30(–75) mm, blade ovate to lanceolate or elliptic, base tapered to cuneate, margins entire or ± repand, denticulate, or dentate, apex obtuse to acute; cauline 2–5 pairs, short-petiolate or sessile, 25–60(–85) × 3–15(–25) mm, blade lanceolate to oblanceolate, base tapered to clasping or cordate-clasping, margins entire, sometimes ± repand to denticulate, apex obtuse to acute. |
basal and cauline, ± leathery or not, glabrous, sometimes sparsely retrorsely hairy along midvein abaxially, not glaucous; basal and proximal cauline (13–)40–90 × (3–)6–18 mm, blade ovate to oblanceolate or lanceolate, base tapered, margins ± serrate or dentate, apex obtuse to acute; cauline 2–6 pairs, short-petiolate or sessile, 15–55 × 3–10 mm, blade lanceolate, proximals sometimes oblanceolate to oblong, base tapered or truncate, margins entire or ± serrate to ± dentate, apex obtuse to acute. |
interrupted or continuous, secund, (2–)5–35 cm, axis sparsely to densely glandular-pubescent, verticillasters 2–5(–7), cymes 2–4-flowered, 2 per node; proximal bracts lanceolate, 11–85 × 1–18 mm, margins entire or ± repand proximally; peduncles and pedicels ascending to erect, glandular-pubescent. |
continuous or ± interrupted, cylindric, 4–19 cm, axis glabrous or ± glandular-pubescent, verticillasters 4–8, cymes (1 or)2–6-flowered, 2 per node; proximal bracts lanceolate, 9–40 × 3–15 mm, margins entire or ± serrate to dentate; peduncles and pedicels erect, sometimes ascending, glandular-pubescent. |
calyx lobes lanceolate, 7–10 × 1–2.2 mm, glandular-pubescent; corolla brownish purple to bluish violet, sometimes ochroleucous, lavender, or white, with alternating white or lavender and purple nectar guides in dark-colored forms, with lavender nectar guides in light-colored forms, ventricose to ventricose-ampliate, 20–27(–30) mm, glandular-pubescent externally, sparsely white-villous internally abaxially, tube 5–8 mm, throat abruptly inflated, 8–10 mm diam., 2-ridged abaxially; stamens included or longer pair reaching orifice, pollen sacs opposite, explanate, 1–1.4 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth; staminode 12–15 mm, reaching orifice or exserted, 0.6–1.1 mm diam., tip straight to slightly recurved, distal 1–3 mm sparsely to densely villous, hairs yellow, to 1 mm; style 12–16 mm. |
calyx lobes ovate, 2.8–4.8 × 0.9–1.8 mm, glandular-pubescent; corolla blue to light blue or violet, with reddish purple nectar guides, funnelform, 13–20 mm, glandular-pubescent externally, sparsely white-villous internally abaxially, tube 4–5 mm, throat gradually inflated, 4–5 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, subexplanate to explanate, 0.5–0.8 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth or papillate; staminode 8–9 mm, included, 0.2–0.3 mm diam., tip straight to recurved, distal 0.5–2 mm ± pilose, hairs golden yellow, to 0.8 mm; style 9–11 mm. |
6–9 × 4–5 mm, glandular-puberulent distally. |
4–6 × 3–4 mm, glabrous. |
= 16. |
= 16. |
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Flowering Jun–Aug(–Sep). |
Flowering May–Aug. |
Rocky slopes in subalpine forests, alpine meadows. |
Gravelly slopes, cliffs, rocky clearings, roadcuts in coniferous forests. |
2100–3700 m. (6900–12100 ft.) |
800–2800 m. (2600–9200 ft.) |
AZ; CO; ID; MT; NM; UT; WY
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ID; MT; AB; BC
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Penstemon whippleanus is known from forests and meadows of the Southern and Central Rocky mountains and westward into the Wasatch Mountains and Utah Plateaus. Plants with light-colored corollas often are found growing next to plants with dark-colored corollas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon caelestinus has been treated as a synonym of P. albertinus (D. D. Keck 1945; A. Cronquist 1959). D. V. Clark (1971) grew these and related species in transplant gardens and considered P. caelestinus to be distinct based on flower color, leaf shape and color, phenology, and habitat. A broad zone of hybridization between P. albertinus and P. wilcoxii exists in east-central Idaho and western Montana, especially in habitats disturbed by humans (Clark). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 227. |
FNA vol. 17, p. 190. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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P. caelestinus |
A. Gray: Proc. Amer. Acad. Arts 6: 73. (1862) — (as Pentstemon) |
Greene: Leafl. Bot. Observ. Crit. 1: 167. (1906) — (as Pentstemon) |
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