Washington beardtongue, Washington penstemon
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Cascade beardtongue, Cascade penstemon, Cascades penstemon, coast penstemon, serrulate penstemon
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Herbs. |
Herbs or subshrubs. |
erect, 3–25 cm, glabrous or ± retrorsely hairy, not glaucous. |
ascending to erect, 13–80 cm, glabrous or retrorsely hairy, hairs pointed, not glaucous. |
basal and cauline, not leathery, glabrous; basal and proximal cauline (10–)17–60 × 5–18 mm, blade spatulate to oblanceolate, lanceolate, or elliptic, base tapered, margins entire, apex rounded to obtuse or acute; cauline 2–6 pairs, short-petiolate or sessile, 10–55 × 3–15 mm, blade ovate to lanceolate, base tapered to clasping, margins entire, apex obtuse to acute. |
cauline, opposite, glabrous, sometimes sparsely puberulent, hairs pointed, adaxially along midvein, not glaucous; cauline 3–6 pairs, petiolate proximally or sessile distally, (12–)30–80 × (7–)15–50 mm, blade ovate-oblong proximally, ovate or lanceolate distally, base truncate to tapered proximally, cordate-clasping distally, margins serrate, apex obtuse to acute. |
interrupted, cylindric, 1–6 cm, axis sparsely glandular-pubescent, sometimes also retrorsely hairy, verticillasters 1–3(or 4), cymes 2–6-flowered, 2 per node; proximal bracts lanceolate, 2–15 × 1–7 mm, margins entire; peduncles and pedicels erect, sparsely to moderately glandular-pubescent. |
continuous or interrupted, cylindric, 2–18(–28) cm, axis retrorsely hairy, verticillasters 1–3(–5), cymes 3–10-flowered, 2 per node; proximal bracts ovate, 13–35(–80) × 6–16(–36) mm; peduncles and pedicels ascending to erect, retrorsely hairy to puberulent. |
calyx lobes lanceolate, 3–4 × 0.8–1.2 mm, glandular-pubescent; corolla dark blue, sometimes whitish, without nectar guides, funnelform, 9–12 mm, glandular-pubescent externally, moderately whitish lanate internally abaxially, tube 3–4 mm, throat slightly inflated, 2–3 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, explanate to subexplanate, 0.5–0.6 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth; staminode 5–6 mm, included, 0.2–0.3 mm diam., tip straight to slightly recurved, distal 0.5–1 mm moderately pilose, hairs golden yellow, to 0.5 mm; style 4–6 mm. |
calyx lobes obovate to lanceolate, 3.5–11 × 1–3 mm, sparsely pubescent and ciliolate along margins, sometimes glabrate; corolla blue to violet or purple, without nectar guides, weakly ventricose, 17–25(–28) mm, glabrous externally, glabrous or sparsely to densely white-lanate internally abaxially, tube 4–6 mm, throat gradually inflated, 5–8 mm diam., 2-ridged abaxially; stamens included, filaments glabrous, pollen sacs parallel, 1.1–1.6 mm, distal 1/2–2/3 indehiscent, sides glabrous, sutures denticulate, teeth to 0.1 mm; staminode 12–14 mm, exserted, 0.3–0.6 mm diam., distal 2–7 mm sparsely lanate, hairs yellow, to 1.5 mm; style 10–16 mm. |
4–7 × 2.5–4 mm, glabrous. |
5–8 × 3–5.5 mm. |
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= 16. |
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Flowering Jul–Aug(–Sep). |
Flowering Jun–Aug(–Sep). |
Alpine meadows, rocky slopes, grassy openings in pine-fir forests. |
Sandy to rocky stream banks, gullies, slopes, roadcuts, coniferous forests. |
1800–2300 m. (5900–7500 ft.) |
0–1800 m. (0–5900 ft.) |
WA
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AK; OR; WA; BC
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Penstemon washingtonensis is known from Chelan and Okanogan counties. Except for its glandular-pubescent inflorescence, P. washingtonensis is morphologically similar to P. procerus var. tolmiei. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon serrulatus occurs from southeastern Alaska through western British Columbia and western Washington to southwestern Oregon, largely in the Cascade and Coast ranges. Herbarium specimens sometimes are misidentified as P. ovatus, which has glandular-pubescent inflorescences and navicular pollen sacs. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 226. |
FNA vol. 17, p. 244. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
D. D. Keck: Amer. Midl. Naturalist 33: 150. (1945) |
Menzies ex Smith: in A. Rees, Cycl. 26: Pentstemon no. 5. (1814) — (as Pentstemon serrulata) |
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