Washington beardtongue, Washington penstemon
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upland beardtongue
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Herbs. |
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erect, 3–25 cm, glabrous or ± retrorsely hairy, not glaucous. |
erect, (10–)30–80 cm, glabrous, not glaucous. |
basal and cauline, not leathery, glabrous; basal and proximal cauline (10–)17–60 × 5–18 mm, blade spatulate to oblanceolate, lanceolate, or elliptic, base tapered, margins entire, apex rounded to obtuse or acute; cauline 2–6 pairs, short-petiolate or sessile, 10–55 × 3–15 mm, blade ovate to lanceolate, base tapered to clasping, margins entire, apex obtuse to acute. |
basal and cauline, not leathery, glabrous, not glaucous; basal and proximal cauline 26–140 × 3–19 mm, blade oblanceolate, base tapered, margins entire, apex obtuse to acute; cauline 2–5 pairs, sessile, 50–90 × (3–)6–9(–18) mm, blade oblanceolate to lanceolate, base clasping, apex acute. |
interrupted, cylindric, 1–6 cm, axis sparsely glandular-pubescent, sometimes also retrorsely hairy, verticillasters 1–3(or 4), cymes 2–6-flowered, 2 per node; proximal bracts lanceolate, 2–15 × 1–7 mm, margins entire; peduncles and pedicels erect, sparsely to moderately glandular-pubescent. |
continuous to ± interrupted, secund or cylindric, 5–40 cm, axis glabrous or ± glandular-pubescent, verticillasters (4 or)5–9, cymes 1–5-flowered, 2 per node; proximal bracts lanceolate, 15–54 × 2–15 mm; peduncles and pedicels glabrous or glandular-pubescent, peduncles to 11 mm, pedicels 1–10 mm. |
calyx lobes lanceolate, 3–4 × 0.8–1.2 mm, glandular-pubescent; corolla dark blue, sometimes whitish, without nectar guides, funnelform, 9–12 mm, glandular-pubescent externally, moderately whitish lanate internally abaxially, tube 3–4 mm, throat slightly inflated, 2–3 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, explanate to subexplanate, 0.5–0.6 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth; staminode 5–6 mm, included, 0.2–0.3 mm diam., tip straight to slightly recurved, distal 0.5–1 mm moderately pilose, hairs golden yellow, to 0.5 mm; style 4–6 mm. |
calyx lobes ovate, 3.5–8 × 1.8–3 mm, margins erose, glabrous or glandular-pubescent; corolla blue to bluish violet, with or without reddish purple nectar guides, ventricose, 17–25(–30) mm, glabrous or glandular-pubescent externally, glabrous or ± glandular-pubescent internally abaxially, tube 5–8 mm, throat gradually inflated, slightly constricted at orifice, 6–8 mm diam., rounded abaxially; stamens: longer pair reaching orifice or slightly exserted, pollen sacs opposite, navicular, 1.2–1.8 mm, dehiscing incompletely, connective not splitting, sides sparsely to moderately hispid or pubescent, hairs whitish, to 0.3 mm, sutures denticulate, teeth to 0.1 mm; staminode 12–15 mm, reaching orifice or slightly exserted, 0.4–0.6 mm diam., tip straight to recurved, distal 3–5 mm sparsely villous, hairs yellow, to 1 mm; style 15–18 mm. |
4–7 × 2.5–4 mm, glabrous. |
8–12 × 4–6 mm. |
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= 16. |
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Flowering Jul–Aug(–Sep). |
Flowering Jun–Aug. |
Alpine meadows, rocky slopes, grassy openings in pine-fir forests. |
Sagebrush shrublands, openings in pine forests. |
1800–2300 m. (5900–7500 ft.) |
2400–3600 m. (7900–11800 ft.) |
WA
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CO; WY |
Penstemon washingtonensis is known from Chelan and Okanogan counties. Except for its glandular-pubescent inflorescence, P. washingtonensis is morphologically similar to P. procerus var. tolmiei. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon saxosorum is known in north-central and northwestern Colorado and south-central Wyoming. Populations in the Bridger Basin in Moffat County, Colorado, and Daggett and Uintah counties, Utah, lie between the main ranges of P. saxosorum and P. subglaber. Plants in those populations have pollen sacs 0.7–0.9 mm; corollas sparsely glandular and 18–20 mm; and corolla throats 5–7 mm in diameter. M. L. Moorman (1982) believed that they represented an undescribed species. Pending further study, Colorado plants are referred to P. saxosorum and Utah plants to P. subglaber. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 226. |
FNA vol. 17, p. 178. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
D. D. Keck: Amer. Midl. Naturalist 33: 150. (1945) |
Pennell: Contr. U.S. Natl. Herb. 20: 349. (1920) |
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