Washington beardtongue, Washington penstemon
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Blue Mountain beardtongue, Blue Mountain penstemon, Blue Mountains beardtongue, Pennell's beardtongue, Pennell's penstemon
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Herbs. |
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erect, 3–25 cm, glabrous or ± retrorsely hairy, not glaucous. |
ascending to erect, 18–85 cm, glabrous, not glaucous. |
basal and cauline, not leathery, glabrous; basal and proximal cauline (10–)17–60 × 5–18 mm, blade spatulate to oblanceolate, lanceolate, or elliptic, base tapered, margins entire, apex rounded to obtuse or acute; cauline 2–6 pairs, short-petiolate or sessile, 10–55 × 3–15 mm, blade ovate to lanceolate, base tapered to clasping, margins entire, apex obtuse to acute. |
basal and cauline, ± leathery or not, glabrous, not glaucous; basal and proximal cauline 55–250 × 7–40 mm, blade lanceolate to elliptic, base tapered, margins entire, apex obtuse to acute; cauline 3–5 pairs, short-petiolate or sessile, 40–83 × 10–36 mm, blade ovate to lanceolate, base tapered to clasping, apex obtuse to acute. |
interrupted, cylindric, 1–6 cm, axis sparsely glandular-pubescent, sometimes also retrorsely hairy, verticillasters 1–3(or 4), cymes 2–6-flowered, 2 per node; proximal bracts lanceolate, 2–15 × 1–7 mm, margins entire; peduncles and pedicels erect, sparsely to moderately glandular-pubescent. |
continuous, cylindric, 4–35 cm, axis glabrous, verticillasters 4–11, cymes 1–6-flowered, 2 per node; proximal bracts ovate to lanceolate, 22–85 × 11–35 mm; peduncles and pedicels glabrous, peduncles to 20 mm, pedicels 1–6 mm. |
calyx lobes lanceolate, 3–4 × 0.8–1.2 mm, glandular-pubescent; corolla dark blue, sometimes whitish, without nectar guides, funnelform, 9–12 mm, glandular-pubescent externally, moderately whitish lanate internally abaxially, tube 3–4 mm, throat slightly inflated, 2–3 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, explanate to subexplanate, 0.5–0.6 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth; staminode 5–6 mm, included, 0.2–0.3 mm diam., tip straight to slightly recurved, distal 0.5–1 mm moderately pilose, hairs golden yellow, to 0.5 mm; style 4–6 mm. |
calyx lobes ovate to lanceolate, (5.3–)6.5–8.5 × 1.6–3 mm, apex acuminate to caudate, glabrous; corolla blue to violet or purple, with reddish purple nectar guides, ventricose, 29–35 mm, glabrous externally, glabrous internally, tube 8–11 mm, throat gradually inflated, not constricted at orifice, 9–10 mm diam., slightly 2-ridged abaxially; stamens: longer pair reaching orifice, pollen sacs divergent to opposite, sigmoid, 2–2.7 mm, dehiscing incompletely, proximal 1/5–1/3 indehiscent, connective not splitting, sides sparsely hirsute, hairs white or tan, to 0.1 mm, especially near connective, sometimes merely scurfy near connective, sutures denticulate, teeth to 0.1 mm; staminode 16–20 mm, reaching orifice, 0.5–1 mm diam., tip straight, distal 4–6 mm ± uniformly sparsely pilose, hairs yellow, to 1 mm; style 20–24 mm. |
4–7 × 2.5–4 mm, glabrous. |
10–13 × 8–10 mm. |
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Flowering Jul–Aug(–Sep). |
Flowering May–Jun. |
Alpine meadows, rocky slopes, grassy openings in pine-fir forests. |
Rocky ridges, sandy to rocky slopes, coniferous forests. |
1800–2300 m. (5900–7500 ft.) |
1300–1800 m. (4300–5900 ft.) |
WA
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OR; WA
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Penstemon washingtonensis is known from Chelan and Okanogan counties. Except for its glandular-pubescent inflorescence, P. washingtonensis is morphologically similar to P. procerus var. tolmiei. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon pennellianus occurs primarily in the Blue Mountains of northeastern Oregon (Grant, Umatilla, Union, and Wallowa counties) and southeastern Washington (Asotin, Columbia, and Garfield counties). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 226. |
FNA vol. 17, p. 176. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
D. D. Keck: Amer. Midl. Naturalist 33: 150. (1945) |
D. D. Keck: Amer. Midl. Naturalist 23: 614. (1940) |
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