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Washington beardtongue, Washington penstemon

Inyo beardtongue

Habit Herbs. Herbs.
Stems

erect, 3–25 cm, glabrous or ± retrorsely hairy, not glaucous.

ascending to erect, 11–48 cm, retrorsely hairy or puberulent, hairs pointed, not glaucous.

Leaves

basal and cauline, not leathery, glabrous;

basal and proximal cauline (10–)17–60 × 5–18 mm, blade spatulate to oblanceolate, lanceolate, or elliptic, base tapered, margins entire, apex rounded to obtuse or acute;

cauline 2–6 pairs, short-petiolate or sessile, 10–55 × 3–15 mm, blade ovate to lanceolate, base tapered to clasping, margins entire, apex obtuse to acute.

basal and cauline, opposite, puberulent, hairs pointed, sometimes glabrate abaxially, not glaucous;

basal and proximal cauline (12–)25–100 × 6–15(–25) mm, blade spatulate to oblanceolate, base tapered, margins entire, apex rounded to obtuse or acute;

cauline 2–4 pairs, sessile or proximals short-petiolate, 24–70 × 7–20 mm, blade elliptic or oblanceolate to lanceolate or oblong, base tapered to cordate-clasping, margins entire, apex obtuse to acute.

Thyrses

interrupted, cylindric, 1–6 cm, axis sparsely glandular-pubescent, sometimes also retrorsely hairy, verticillasters 1–3(or 4), cymes 2–6-flowered, 2 per node;

proximal bracts lanceolate, 2–15 × 1–7 mm, margins entire;

peduncles and pedicels erect, sparsely to moderately glandular-pubescent.

interrupted, cylindric, 8–20 cm, axis glandular-pubescent, verticillasters 3–7, cymes 1–4-flowered, 2 per node;

proximal bracts lanceolate, 13–45 × 4–12 mm;

peduncles and pedicels erect, glandular-pubescent.

Flowers

calyx lobes lanceolate, 3–4 × 0.8–1.2 mm, glandular-pubescent;

corolla dark blue, sometimes whitish, without nectar guides, funnelform, 9–12 mm, glandular-pubescent externally, moderately whitish lanate internally abaxially, tube 3–4 mm, throat slightly inflated, 2–3 mm diam., 2-ridged abaxially;

stamens included, pollen sacs opposite, explanate to subexplanate, 0.5–0.6 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth;

staminode 5–6 mm, included, 0.2–0.3 mm diam., tip straight to slightly recurved, distal 0.5–1 mm moderately pilose, hairs golden yellow, to 0.5 mm;

style 4–6 mm.

calyx lobes lanceolate, 8–10 × 1.4–2.6 mm, glandular-pubescent;

corolla blue to lavender or violet, without nectar guides, ampliate, 24–30 mm, glandular-pubescent externally, glabrous internally, tube 8–10 mm, throat gradually inflated, 6–8 mm diam., 2-ridged abaxially;

stamens included, filaments glabrous, pollen sacs parallel, 1.6–1.9 mm, distal 1/3 indehiscent, sides glabrous, sutures denticulate, teeth to 0.1 mm;

staminode 13–15 mm, included, 0.4–0.5 mm diam., distal 5–8 mm densely pilose, hairs yellow, to 1.3 mm;

style 22–25 mm.

Capsules

4–7 × 2.5–4 mm, glabrous.

7–11 × 5–6 mm.

Penstemon washingtonensis

Penstemon papillatus

Phenology Flowering Jul–Aug(–Sep). Flowering Jun–Aug.
Habitat Alpine meadows, rocky slopes, grassy openings in pine-fir forests. Rocky openings, pine-juniper woodlands, pine forests.
Elevation 1800–2300 m. (5900–7500 ft.) 2000–2900 m. (6600–9500 ft.)
Distribution
from FNA
WA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Penstemon washingtonensis is known from Chelan and Okanogan counties. Except for its glandular-pubescent inflorescence, P. washingtonensis is morphologically similar to P. procerus var. tolmiei.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon papillatus is limited to the High Sierra Nevada and Inyo and White mountains in Inyo and Mono counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 226. FNA vol. 17, p. 239.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon Plantaginaceae > Penstemon > subg. Penstemon > sect. Saccanthera
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Name authority D. D. Keck: Amer. Midl. Naturalist 33: 150. (1945) J. T. Howell: Leafl. W. Bot. 2: 119. (1938)
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