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Heller's beardtongue

Arkansas beardtongue

Habit Herbs.
Stems

erect, 30–65 cm, retrorsely hairy.

ascending to erect, (18–)25–45(–61) cm, retrorsely hairy, sometimes also ± glandular-pubescent distally, not glaucous.

Leaves

basal and cauline or, sometimes, basal absent or withering, not leathery to ± leathery, glabrous or retrorsely hairy along midvein and, sometimes, along margins;

basal and proximal cauline petiolate, 22–120(–190) × 7–40(–65) mm, blade oblanceolate to elliptic, base tapered, margins entire or serrate to dentate, apex obtuse to acute;

cauline 6–8(–10) pairs, sessile, 30–95(–115) × 7–30 mm, blade ovate to lanceolate to linear, base tapered to cordate clasping, margins entire or serrate, apex acute.

basal and cauline, basal often withering by anthesis, not leathery, glabrous or sparsely retrorsely hairy, especially along midvein and margins, abaxially, glabrous or sparsely puberulent adaxially;

basal and proximal cauline 20–95 × 5–28 mm, blade spatulate or obovate to ovate, base tapered, margins subentire to ± serrate or dentate, apex rounded to obtuse or acute;

cauline (6–)8–12 pairs, short-petiolate or sessile, 22–80 × 4–20 mm, blade lanceolate, base tapered to clasping, margins ± serrate or dentate, apex acute to acuminate.

Thyrses

interrupted, cylindric, 5–20 cm, axis glandular-pubescent, verticillasters (3–)4–6, cymes 2–4(–6)-flowered, 2 per node;

proximal bracts ovate to lanceolate, 14–50 × 6–17 mm;

peduncles and pedicels glandular-pubescent.

interrupted, conic, 6–24 cm, axis retrorsely hairy and ± glandular-pubescent, verticillasters 4–8, cymes 3–13-flowered, 2 per node;

proximal bracts lanceolate, 6–40 × 2–12 mm, margins entire or ± dentate;

peduncles and pedicels ascending, retrorsely hairy and ± glandular-pubescent.

Flowers

calyx lobes ovate to lanceolate, 6.5–13 × 2.5–4.8 mm, glandular-pubescent;

corolla pink to pinkish red or reddish violet, with red or crimson nectar guides, ventricose, 22–35 mm, glandular-pubescent internally, tube 8–11 mm, throat gradually inflated, not constricted at orifice, 10–13 mm diam., rounded abaxially;

stamens: longer pair exserted, pollen sacs opposite, explanate, 0.9–1.2 mm, dehiscing completely, sutures smooth;

staminode 17–20 mm, included or reaching orifice, 0.4–0.6 mm diam., tip straight to recurved, distal 8–12 mm sparsely to moderately pilose, hairs yellow, to 1 mm;

style 18–24 mm.

calyx lobes ovate, 2.8–4.5 × 1.1–2 mm, sparsely glandular-pubescent;

corolla white, with reddish purple nectar guides, tubular to tubular-funnelform, 15–19 mm, glandular-pubescent externally, moderately whitish lanate internally abaxially, tube 4–6 mm, throat slightly inflated, 4–7 mm diam., 2-ridged abaxially;

stamens included, pollen sacs opposite, navicular, 0.8–1 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate;

staminode 10–12 mm, exserted, 0.2–0.4 mm diam., tip straight to slightly recurved, distal 5–7 mm ± villous, hairs yellow or golden yellow, to 1.2 mm;

style 9–11 mm.

Capsules

(7–)11–14 × 5–8.5 mm.

4–7 × 4–5 mm, glabrous.

2n

= 16.

Penstemon triflorus

Penstemon arkansanus

Phenology Flowering Apr–Jun.
Habitat Rocky oak-hickory woodlands, glades, rocky stream bottoms.
Elevation 60–300 m. (200–1000 ft.)
Distribution
from FNA
TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AR; MO; OK
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Penstemon triflorus occurs largely on the Edwards Plateau, though a few populations are known from adjacent physiographic provinces in central Texas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon arkansanus is centered in the Interior Highlands. The species is most similar to P. pallidus, from which it differs by its narrower, less hairy, and more numerous cauline leaves. Some specimens from the narrow zone of sympatry in north-central Arkansas and south-central Missouri exhibit varying degrees of morphologic intermediacy. Penstemon arkansanus has been reported from Illinois (R. H. Mohlenbrock 1986) and Texas (S. D. Jones et al. 1997); those reports seem unlikely.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Cauline leaf blades ovate to lanceolate, margins sharply serrate; calyx lobes 8–13 × 3–4.8 mm; corollas 30–35 mm.
var. triflorus
1. Cauline leaf blades lanceolate to linear, margins entire or obscurely, rarely sharply, serrate; calyx lobes 6.5–8.5 × 2.4–3.5 mm; corollas 22–30 mm.
var. integrifolius
Source FNA vol. 17, p. 142. FNA vol. 17, p. 193.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Cristati Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Subordinate taxa
P. triflorus var. integrifolius, P. triflorus var. triflorus
Synonyms P. multicaulis, P. pallidus subsp. arkansanus, P. wherryi
Name authority A. Heller: Contr. Herb. Franklin Marshall Coll. 1: 92, plate 8. (1895) — (as Pentstemon) Pennell: Proc. Acad. Nat. Sci. Philadelphia 73: 493. (1922)
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