Heller's beardtongue
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Siskiyou beardtongue, tongue leaf penstemon
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Herbs. |
erect, 30–65 cm, retrorsely hairy. |
ascending to erect, (10–)30–80 cm, glabrous, not glaucous. |
basal and cauline or, sometimes, basal absent or withering, not leathery to ± leathery, glabrous or retrorsely hairy along midvein and, sometimes, along margins; basal and proximal cauline petiolate, 22–120(–190) × 7–40(–65) mm, blade oblanceolate to elliptic, base tapered, margins entire or serrate to dentate, apex obtuse to acute; cauline 6–8(–10) pairs, sessile, 30–95(–115) × 7–30 mm, blade ovate to lanceolate to linear, base tapered to cordate clasping, margins entire or serrate, apex acute. |
basal and cauline, not leathery, glabrous; basal and proximal cauline 30–150 × 6–40 mm, blade ovate to elliptic or oval, base tapered, margins serrate to dentate, sometimes denticulate or entire, apex obtuse to acute; cauline 2–4 pairs, sessile or proximals short-petiolate, 11–90 × 5–50 mm, blade oblanceolate to oblong or triangular-ovate, base tapered to cordate-clasping, margins serrate to denticulate, rarely entire, apex obtuse to acute. |
interrupted, cylindric, 5–20 cm, axis glandular-pubescent, verticillasters (3–)4–6, cymes 2–4(–6)-flowered, 2 per node; proximal bracts ovate to lanceolate, 14–50 × 6–17 mm; peduncles and pedicels glandular-pubescent. |
interrupted, sometimes continuous, cylindric, (4–)8–40 cm, axis glandular-pubescent, verticillasters 3–10, cymes (2 or)3–15-flowered, 2 per node; proximal bracts ovate to triangular, 10–85 × 7–40 mm, margins serrate or entire; peduncles and pedicels ascending, glandular-pubescent. |
calyx lobes ovate to lanceolate, 6.5–13 × 2.5–4.8 mm, glandular-pubescent; corolla pink to pinkish red or reddish violet, with red or crimson nectar guides, ventricose, 22–35 mm, glandular-pubescent internally, tube 8–11 mm, throat gradually inflated, not constricted at orifice, 10–13 mm diam., rounded abaxially; stamens: longer pair exserted, pollen sacs opposite, explanate, 0.9–1.2 mm, dehiscing completely, sutures smooth; staminode 17–20 mm, included or reaching orifice, 0.4–0.6 mm diam., tip straight to recurved, distal 8–12 mm sparsely to moderately pilose, hairs yellow, to 1 mm; style 18–24 mm. |
calyx lobes lanceolate, 4–6.5 × 0.9–2 mm, glandular-pubescent; corolla light blue to lavender, purple, or violet, with violet nectar guides, ampliate, 13–18 mm, glandular-pubescent externally, glabrous or sparsely white-lanate internally abaxially, tube 5–7 mm, throat gradually inflated, 4–6 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, navicular, 0.8–1.2 mm, dehiscing completely, connective splitting, sides glabrous, sutures papillate; staminode 10–13 mm, exserted, 0.2–0.4 mm diam., tip straight to slightly recurved, distal 2–6 mm glabrous or sparsely pilose, hairs yellow, to 0.6 mm; style 10–13 mm. |
(7–)11–14 × 5–8.5 mm. |
5–7 × 3–4 mm, glabrous. |
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= 16. |
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Flowering Jun–Aug. |
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Rocky, clayey, or humus soils, openings in coniferous forests. |
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600–2000 m. (2000–6600 ft.) |
TX; n Mexico
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CA; OR
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Varieties 2 (2 in the flora). Penstemon triflorus occurs largely on the Edwards Plateau, though a few populations are known from adjacent physiographic provinces in central Texas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon anguineus occurs primarily in the Cascade, Klamath, and North Coast ranges from southwestern Oregon (Curry, Douglas, Jackson, Josephine, and Klamath counties) south into northwestern California (Del Norte, El Dorado, Glenn, Humboldt, Mendocino, Siskiyou, Tehama, and Trinity counties). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
1. Cauline leaf blades ovate to lanceolate, margins sharply serrate; calyx lobes 8–13 × 3–4.8 mm; corollas 30–35 mm. | var. triflorus |
1. Cauline leaf blades lanceolate to linear, margins entire or obscurely, rarely sharply, serrate; calyx lobes 6.5–8.5 × 2.4–3.5 mm; corollas 22–30 mm. | var. integrifolius |
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FNA vol. 17, p. 142. |
FNA vol. 17, p. 191. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Cristati |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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A. Heller: Contr. Herb. Franklin Marshall Coll. 1: 92, plate 8. (1895) — (as Pentstemon) |
Eastwood: Bull. Torrey Bot. Club 32: 208. (1905) — (as Pentstemon) |
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