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Arizona scarlet-bugler, hackberry beardtongue

Tidestrom's beardtongue

Stems

ascending to erect, 20–60 cm, glaucous, sometimes not.

ascending to erect, 25–50 cm, retrorsely hairy, not glaucous.

Leaves

glabrous, glaucous;

basal and proximal cauline 28–100 × 2–11 mm, blade oblanceolate to elliptic, base tapered, margins entire, apex obtuse to acute;

cauline 3–6 pairs, sessile, (11–)30–80(–120) × 2–9 mm, blade lanceolate to linear or linear-subulate, base clasping to auriculate-clasping, margins entire, apex rounded to acute.

basal and cauline, ± leathery or not, retrorsely hairy, not glaucous;

basal and proximal cauline 40–120 × 4–33 mm, blade obovate to oblanceolate, base tapered, margins entire, apex obtuse to acute;

cauline 3–5 pairs, short-petiolate or sessile, 35–70(–90) × 8–22 mm, blade lanceolate to oblanceolate, base tapered to truncate or clasping, apex obtuse to acute.

Thyrses

interrupted, cylindric, 6–26 cm, axis glabrous, verticillasters 5–9, cymes 1–4-flowered;

proximal bracts lanceolate to linear, 6–33 × 1–4 mm;

peduncles and pedicels erect, glabrous.

interrupted, secund, 9–22 cm, axis glabrous or sparsely retrorsely hairy, verticillasters 7–15, cymes 5- or 6-flowered, 2 per node;

proximal bracts lanceolate, 12–50 × 3–16 mm;

peduncles and pedicels glabrous or sparsely retrorsely hairy.

Flowers

calyx lobes ovate to lanceolate, 2–4.5 × 1.4–2.1 mm, margins entire or erose, glabrous or ± glandular along margins;

corolla scarlet, without nectar guides, nearly radially symmetric, weakly bilabiate, salverform, 18–26 mm, glabrous externally, glabrous internally, tube 4–7 mm, 1.6–3 times as long as calyx lobes, throat slightly inflated, 3–5 mm diam., rounded abaxially;

stamens: longer pair reaching orifice, pollen sacs explanate, 0.7–0.9 mm, sutures smooth;

staminode 13–14 mm, essentially terete, 0.1 mm diam., tip straight, glabrous, rarely distal 1–2 mm sparsely papillate, papillae yellow, to 0.2 mm;

style 10–19 mm.

calyx lobes ovate, 2.5–4 × 1.5–1.7(–2) mm, margins narrowly scarious, apex obtuse, sometimes acute to cuspidate, glabrous;

corolla blue to violet, without nectar guides, ventricose, 13–21 mm, glabrous externally, glabrous internally, tube 5–6 mm, throat gradually inflated, not constricted at orifice, 5–7 mm diam., rounded abaxially;

stamens included or longer pair reaching orifice, pollen sacs divergent, navicular, 1.3–1.7 mm, dehiscing incompletely, proximal 1/3–1/2 sometimes indehiscent, connective not splitting, sides glabrous, sutures denticulate, teeth to 0.1 mm;

staminode 8–11 mm, included, 0.6–0.8 mm diam., tip straight, distal 5–8 mm sparsely to moderately pilose, hairs yellow, to 1 mm;

style 11–13 mm.

Capsules

6–8 × 4–5 mm.

6–9 × 4.5–6.5 mm.

2n

= 16.

Penstemon subulatus

Penstemon tidestromii

Phenology Flowering (Feb–)Mar–May. Flowering May–Jul.
Habitat Rocky slopes, mesas, canyons. Sagebrush shrublands, pinyon-juniper woodlands.
Elevation 500–1300 m. (1600–4300 ft.) 1600–2500 m. (5200–8200 ft.)
Distribution
from FNA
AZ
[WildflowerSearch map]
[BONAP county map]
from FNA
UT
[BONAP county map]
Discussion

Penstemon subulatus has been documented in Gila, Graham, Maricopa, Mohave, Pinal, and Yuma counties in central Arizona.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon tidestromii is known from Juab and Sanpete counties. Specimens from the San Pitch Mountains with retrorsely hairy stems, corollas obscurely glandular, and pollen sacs 1.5–1.7 mm and sparsely hispid might be hybrids or introgressants between P. subglaber and P. tidestromii. Penstemon leptanthus, known only from the type collection (Ward 280, US), seems to be morphologically confluent with P. tidestromii.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 154. FNA vol. 17, p. 181.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Gentianoides Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Synonyms P. leptanthus
Name authority M. E. Jones: Contr. W. Bot. 12: 63. (1908) — (as Pentstemon) Pennell: Contr. U.S. Natl. Herb. 20: 379. (1920)
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