Arizona scarlet-bugler, hackberry beardtongue
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gentian beardtongue
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Herbs. |
ascending to erect, 20–60 cm, glaucous, sometimes not. |
glabrous, glaucous, sometimes not. |
glabrous, glaucous; basal and proximal cauline 28–100 × 2–11 mm, blade oblanceolate to elliptic, base tapered, margins entire, apex obtuse to acute; cauline 3–6 pairs, sessile, (11–)30–80(–120) × 2–9 mm, blade lanceolate to linear or linear-subulate, base clasping to auriculate-clasping, margins entire, apex rounded to acute. |
basal and cauline, sometimes basal few (P. subulatus), or essentially cauline (P. centranthifolius), opposite, leathery, rarely not (P. alamosensis, P. parryi), glabrous, rarely scabrous, glaucous, rarely not; basal and proximal cauline petiolate; cauline sessile or short-petiolate, blade ovate, oblanceolate, lanceolate, oblong, elliptic, linear, or linear-subulate, margins entire, rarely toothed. |
interrupted, cylindric, 6–26 cm, axis glabrous, verticillasters 5–9, cymes 1–4-flowered; proximal bracts lanceolate to linear, 6–33 × 1–4 mm; peduncles and pedicels erect, glabrous. |
± interrupted, cylindric or ± secund, axis glabrous, rarely scabrous, cymes 2 per node, sometimes 1 per node (P. alamosensis); peduncles and pedicels spreading to ascending or erect. |
calyx lobes ovate to lanceolate, 2–4.5 × 1.4–2.1 mm, margins entire or erose, glabrous or ± glandular along margins; corolla scarlet, without nectar guides, nearly radially symmetric, weakly bilabiate, salverform, 18–26 mm, glabrous externally, glabrous internally, tube 4–7 mm, 1.6–3 times as long as calyx lobes, throat slightly inflated, 3–5 mm diam., rounded abaxially; stamens: longer pair reaching orifice, pollen sacs explanate, 0.7–0.9 mm, sutures smooth; staminode 13–14 mm, essentially terete, 0.1 mm diam., tip straight, glabrous, rarely distal 1–2 mm sparsely papillate, papillae yellow, to 0.2 mm; style 10–19 mm. |
calyx lobes: entire or erose, ± scarious, glabrous or sparsely glandular or glandular-pubescent; corolla red, scarlet, crimson, magenta, pink, orange, lavender, violet, reddish violet, or purple, bilaterally symmetric or nearly radially symmetric, bilabiate or weakly bilabiate, not personate, salverform, tubular-salverform, or tubular-funnelform, sometimes ventricose, glabrous or glandular-pubescent to glandular externally, glabrous or glandular-pubescent internally abaxially or wholly, throat slightly to gradually inflated, not constricted at orifice, rounded or slightly 2-ridged abaxially; stamens included or longer pair reaching orifice, filaments glabrous, pollen sacs opposite, dehiscing completely, explanate, sometimes navicular, connective splitting, sides glabrous, sutures smooth, sometimes papillate; staminode included, flattened distally or ± terete, 0.1–1(–2.5) mm diam., tip straight or recurved, glabrous or distal 10–40(–70)% papillate or hairy, papillae or hairs to 1.5 mm; style glabrous. |
6–8 × 4–5 mm. |
glabrous. |
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brown to black, angled, 1–4 mm. |
= 16. |
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Flowering (Feb–)Mar–May. |
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Rocky slopes, mesas, canyons. |
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500–1300 m. (1600–4300 ft.) |
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AZ
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w United States; n Mexico |
Penstemon subulatus has been documented in Gila, Graham, Maricopa, Mohave, Pinal, and Yuma counties in central Arizona. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 11 (10 in the flora). Section Gentianoides is not monophyletic. Molecular data (C. A. Wessinger et al. 2016) placed some of the species in a clade that includes sect. Coerulei and other species in a clade that includes sect. Spectabiles; still others may be allied with sect. Glabri (A. D. Wolfe et al. 2006). D. D. Keck (1937) included most of the species of sect. Gentianoides in subsect. Centranthifolii [an unpublished name] of sect. Peltanthera D. D. Keck, including Penstemon cerrosensis Kellogg, known only from Cedros Island off the west coast of Baja California, Mexico. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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1. Corollas glabrous externally. | → 2 |
2. Corollas salverform, weakly bilabiate. | → 3 |
3. Corollas 25–33 mm, tubes 8–10 mm; cauline leaf blades ovate to lanceolate. | P. centranthifolius |
3. Corollas 18–26 mm, tubes 4–7 mm; cauline leaf blades lanceolate to linear or linear-subulate. | P. subulatus |
2. Corollas tubular-funnelform, bilabiate. | → 4 |
4. Corollas glabrous or glandular-pubescent internally; peduncles and pedicels ascending to erect. | P. confusus |
4. Corollas glabrous internally; peduncles and pedicels spreading to ascending. | P. patens |
1. Corollas glandular-pubescent externally. | → 5 |
5. Corollas white-pilose and glandular-pubescent internally abaxially. | P. parryi |
5. Corollas glandular-pubescent or glabrous internally abaxially. | → 6 |
6. Calyx lobes sparsely glandular-pubescent. | → 7 |
| → 8 |
8. Staminodes 7–9 mm; leaves not leathery. | P. alamosensis |
8. Staminodes 14–20 mm; leaves leathery. | P. havardii |
7. Staminodes retrorsely hairy or sparsely to densely pilose distally. | → 9 |
9. Corollas nearly radially symmetric, weakly bilabiate; staminodes: distal 2–3 mm retrorsely hairy, hairs yellow or whitish, to 1.5 mm. | P. superbus |
9. Corollas bilaterally symmetric, bilabiate; staminodes: distal 2–7 mm sparsely to densely pilose, hairs yellow, to 1.2 mm. | P. wrightii |
| → 10 |
10. Staminodes: distal 2–3 mm retrorsely hairy, hairs yellow or whitish, to 1.5 mm; corollas nearly radially symmetric, weakly bilabiate. | P. superbus |
10. Staminodes glabrous or distal 1–3 mm papillate, papillae golden yellow or reddish yellow, to 0.2 mm; corollas bilaterally symmetric, bilabiate. | → 11 |
11. Corollas red to crimson, tubular-salverform, glandular-pubescent externally and internally; pollen sacs explanate. | P. utahensis |
11. Corollas violet or reddish violet to purple or lavender, tubular-funnelform, glabrous or glandular-pubescent externally, glabrous or glandular-pubescent internally; pollen sacs navicular, sometimes explanate. | → 12 |
12. Corollas glabrous or glandular-pubescent internally; pedicels ascending to erect. | P. confusus |
12. Corollas glabrous internally; pedicels spreading to ascending. | P. patens |
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FNA vol. 17, p. 154. |
FNA vol. 17, p. 150. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Gentianoides |
Plantaginaceae > Penstemon > subg. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
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M. E. Jones: Contr. W. Bot. 12: 63. (1908) — (as Pentstemon) |
G. Don: Gen. Hist. 4: 640. (1837) — (as Pentstemon) |
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