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Arizona scarlet-bugler, hackberry beardtongue

long-sepal beardtongue

Habit Herbs.
Stems

ascending to erect, 20–60 cm, glaucous, sometimes not.

erect, (40–)60–120 cm, puberulent, at least proximally, slightly glaucous or not.

Leaves

glabrous, glaucous;

basal and proximal cauline 28–100 × 2–11 mm, blade oblanceolate to elliptic, base tapered, margins entire, apex obtuse to acute;

cauline 3–6 pairs, sessile, (11–)30–80(–120) × 2–9 mm, blade lanceolate to linear or linear-subulate, base clasping to auriculate-clasping, margins entire, apex rounded to acute.

basal and cauline, basal often withering by anthesis, not leathery, glabrous;

basal and proximal cauline 47–110 × 10–24 mm, blade oblanceolate, base tapered, margins entire or ± serrate, apex rounded to obtuse;

cauline 3–7 pairs, sessile or proximals short-petiolate, (14–)44–155 × (2–)13–46 mm, blade lanceolate to ovate, proximals sometimes oblanceolate, base clasping or tapered, margins serrate, apex acute to acuminate.

Thyrses

interrupted, cylindric, 6–26 cm, axis glabrous, verticillasters 5–9, cymes 1–4-flowered;

proximal bracts lanceolate to linear, 6–33 × 1–4 mm;

peduncles and pedicels erect, glabrous.

interrupted, conic, (5–)14–20 cm, axis sparsely puberulent and glandular-pubescent, rarely glabrous, verticillasters 2–5, cymes (1–)5–15-flowered, 2 per node;

proximal bracts ovate to lanceolate, sometimes linear, 13–80 × (2–)5–32 mm, margins serrate, sometimes entire;

peduncles and pedicels spreading to ascending, sparsely glandular-pubescent.

Flowers

calyx lobes ovate to lanceolate, 2–4.5 × 1.4–2.1 mm, margins entire or erose, glabrous or ± glandular along margins;

corolla scarlet, without nectar guides, nearly radially symmetric, weakly bilabiate, salverform, 18–26 mm, glabrous externally, glabrous internally, tube 4–7 mm, 1.6–3 times as long as calyx lobes, throat slightly inflated, 3–5 mm diam., rounded abaxially;

stamens: longer pair reaching orifice, pollen sacs explanate, 0.7–0.9 mm, sutures smooth;

staminode 13–14 mm, essentially terete, 0.1 mm diam., tip straight, glabrous, rarely distal 1–2 mm sparsely papillate, papillae yellow, to 0.2 mm;

style 10–19 mm.

calyx lobes lanceolate, 5–9 × 0.8–1.9 mm, apex acuminate, sparsely glandular-pubescent;

corolla pale lavender to violet, with faint violet nectar guides, ventricose, 20–35 mm, glandular-pubescent externally, ± white-pubescent internally abaxially, tube 5–7 mm, throat abruptly inflated, 8–11 mm diam., slightly 2-ridged abaxially;

stamens included or longer pair reaching orifice, pollen sacs opposite, navicular, 1.2–1.4 mm, dehiscing completely, connective splitting, sides glabrous, rarely sparsely pubescent, hairs white, to 0.2 mm, sutures papillate;

staminode 11–13 mm, reaching orifice, 0.2–0.3 mm diam., tip straight to slightly recurved, distal 2–5 mm sparsely pubescent, hairs yellowish, to 1 mm;

style 14–16 mm.

Capsules

6–8 × 4–5 mm.

7–8 × 4–5 mm, glabrous.

2n

= 16.

= 96.

Penstemon subulatus

Penstemon calycosus

Phenology Flowering (Feb–)Mar–May. Flowering May–Jun.
Habitat Rocky slopes, mesas, canyons. Woodlands, meadows, rocky slopes, stream banks.
Elevation 500–1300 m. (1600–4300 ft.) 10–300 m. (0–1000 ft.)
Distribution
from FNA
AZ
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; CT; DC; GA; IL; IN; KY; MA; MD; ME; MI; MN; MO; NC; NJ; NY; OH; PA; RI; SC; TN; VA; VT; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

Penstemon subulatus has been documented in Gila, Graham, Maricopa, Mohave, Pinal, and Yuma counties in central Arizona.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon calycosus is concentrated in the Central Lowlands and Interior Low Plateaus; occurrences farther south and east, especially in New England, may be from introductions (F. W. Pennell 1935). The range lies almost entirely within that of P. digitalis, but historic ranges of both P. calycosus and P. digitalis are difficult to delimit due to both past and continuing introduction and spread of both species. Calyx lobe lengths to 12 mm are reported in the literature; those appear to be from fruiting calyces. Penstemon calycosus usually has glabrous anthers, but many specimens, especially from Illinois and Ohio, bear hairs on the adaxial surfaces of the anthers. Some populations from this region exhibit variation in vegetative (leaf size and bract shape and margins) and floral characters (calyx lobe shape and size, corolla color, and anther pubescence), suggesting possible hybridization with P. digitalis, an observation that would be consistent with the findings of A. C. Koelling (1964).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 154. FNA vol. 17, p. 197.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Gentianoides Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Name authority M. E. Jones: Contr. W. Bot. 12: 63. (1908) — (as Pentstemon) Small: Bull. Torrey Bot. Club 25: 470. (1898) — (as Pentstemon)
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