Arizona scarlet-bugler, hackberry beardtongue
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Wilcox's beardtongue, Wilcox's penstemon
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Herbs. |
ascending to erect, 20–60 cm, glaucous, sometimes not. |
ascending to erect, (20–)35–100 cm, glabrous or sparsely to densely puberulent, sometimes also sparsely glandular-pubescent, especially distally, not glaucous. |
glabrous, glaucous; basal and proximal cauline 28–100 × 2–11 mm, blade oblanceolate to elliptic, base tapered, margins entire, apex obtuse to acute; cauline 3–6 pairs, sessile, (11–)30–80(–120) × 2–9 mm, blade lanceolate to linear or linear-subulate, base clasping to auriculate-clasping, margins entire, apex rounded to acute. |
basal and cauline, ± leathery or not, glabrous or puberulent to pubescent, especially along midveins; basal and proximal cauline (25–)35–190 × (5–)12–55 mm, blade triangular to ovate or lanceolate, base tapered, rarely truncate, margins ± serrate or dentate, apex acute, rarely obtuse; cauline 2–5 pairs, short-petiolate or sessile, 35–90 × (8–)15–50 mm, blade ovate to oblong or lanceolate, base tapered to clasping or cordate-clasping, margins serrate to dentate, apex obtuse to acute. |
interrupted, cylindric, 6–26 cm, axis glabrous, verticillasters 5–9, cymes 1–4-flowered; proximal bracts lanceolate to linear, 6–33 × 1–4 mm; peduncles and pedicels erect, glabrous. |
interrupted, cylindric to conic, (5–)8–30 cm, axis sparsely to moderately glandular-pubescent, verticillasters 3–7, cymes 3–11-flowered, 2 per node; proximal bracts ovate to lanceolate, (8–)18–75 × (5–)8–35 mm, margins ± serrate to dentate; peduncles and pedicels spreading to ascending, ± glandular-pubescent. |
calyx lobes ovate to lanceolate, 2–4.5 × 1.4–2.1 mm, margins entire or erose, glabrous or ± glandular along margins; corolla scarlet, without nectar guides, nearly radially symmetric, weakly bilabiate, salverform, 18–26 mm, glabrous externally, glabrous internally, tube 4–7 mm, 1.6–3 times as long as calyx lobes, throat slightly inflated, 3–5 mm diam., rounded abaxially; stamens: longer pair reaching orifice, pollen sacs explanate, 0.7–0.9 mm, sutures smooth; staminode 13–14 mm, essentially terete, 0.1 mm diam., tip straight, glabrous, rarely distal 1–2 mm sparsely papillate, papillae yellow, to 0.2 mm; style 10–19 mm. |
calyx lobes ovate, 2.4–5.5 × 0.9–1.8 mm, margins broadly scarious, glandular-pubescent; corolla blue to violet or light blue, with reddish purple nectar guides, funnelform, 13–23 mm, glandular-pubescent externally, sparsely white- or yellow-villous internally abaxially, tube 4–8 mm, throat gradually inflated, 4–9 mm diam., 2-ridged abaxially; stamens included, pollen sacs opposite, navicular to subexplanate, 0.8–1 mm, dehiscing completely, connective splitting, sides glabrous, sutures smooth or ± papillate; staminode 8–11 mm, reaching orifice, 0.2–0.3 mm diam., tip straight to recurved, distal 1–5 mm pilose to villous, hairs yellow, to 1.2 mm; style 9–13 mm. |
6–8 × 4–5 mm. |
4–6.5 × 3–4 mm, glabrous. |
= 16. |
= 16, 32. |
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Flowering (Feb–)Mar–May. |
Flowering May–Aug. |
Rocky slopes, mesas, canyons. |
Rocky or gravelly slopes in forests and woodlands, roadcuts. |
500–1300 m. (1600–4300 ft.) |
300–2300 m. (1000–7500 ft.) |
AZ
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ID; MT; OR; WA
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Penstemon subulatus has been documented in Gila, Graham, Maricopa, Mohave, Pinal, and Yuma counties in central Arizona. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Penstemon wilcoxii hybridizes with P. albertinus in areas of sympatry in east-central Idaho and west-central Montana (D. D. Keck 1945; A. Cronquist 1959; D. V. Clark 1971). Putative hybrids and/or introgressants exhibit varying degrees of morphologic intermediacy between the parent species. Many herbarium specimens are difficult to determine with confidence. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
FNA vol. 17, p. 154. |
FNA vol. 17, p. 227. |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Gentianoides |
Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis |
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wrightii, P. xylus, P. yampaënsis |
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P. ovatus var. pinetorum |
M. E. Jones: Contr. W. Bot. 12: 63. (1908) — (as Pentstemon) |
Rydberg: Bull. Torrey Bot. Club 28: 28. (1901) — (as Pentstemon) |
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