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Mancos penstemon, stiff beardtongue, stiff penstemon

long-sepal beardtongue

Habit Herbs.
Stems

ascending to erect, (11–)16–55(–70) cm, glabrous, not glaucous.

erect, (40–)60–120 cm, puberulent, at least proximally, slightly glaucous or not.

Leaves

basal and cauline, ± leathery or not, glabrous, not glaucous;

basal and proximal cauline 60–90(–150) × 5–12 mm, blade oblanceolate to lanceolate, base tapered, margins entire, apex acute;

cauline 3–5 pairs, sessile, 18–80 × 3–18 mm, blade lanceolate, base tapered to truncate or clasping, apex acute.

basal and cauline, basal often withering by anthesis, not leathery, glabrous;

basal and proximal cauline 47–110 × 10–24 mm, blade oblanceolate, base tapered, margins entire or ± serrate, apex rounded to obtuse;

cauline 3–7 pairs, sessile or proximals short-petiolate, (14–)44–155 × (2–)13–46 mm, blade lanceolate to ovate, proximals sometimes oblanceolate, base clasping or tapered, margins serrate, apex acute to acuminate.

Thyrses

continuous or interrupted, ± secund, 5–40 cm, axis glabrous, verticillasters 4–8(–12), cymes 2- or 3(–5)-flowered, 2 per node;

proximal bracts lanceolate, 12–46(–115) × 2–15 mm, apex acuminate;

peduncles and pedicels glabrous or ± glandular, peduncles to 20 mm, pedicels 2–13(–18) mm.

interrupted, conic, (5–)14–20 cm, axis sparsely puberulent and glandular-pubescent, rarely glabrous, verticillasters 2–5, cymes (1–)5–15-flowered, 2 per node;

proximal bracts ovate to lanceolate, sometimes linear, 13–80 × (2–)5–32 mm, margins serrate, sometimes entire;

peduncles and pedicels spreading to ascending, sparsely glandular-pubescent.

Flowers

calyx lobes ovate to lanceolate, (4.5–)6–8(–9) × 2–3 mm, apex acuminate to caudate, glabrous or ± glandular proximally;

corolla lavender to blue, without nectar guides, ventricose, (19–)25–32 mm, glabrous externally, glabrous internally, tube 5–7 mm, throat gradually to abruptly inflated, not constricted at orifice, 7–10 mm diam., slightly 2-ridged abaxially;

stamens: longer pair slightly exserted, pollen sacs divergent, navicular-sigmoid, 1.3–2.3 mm, dehiscing incompletely, proximal 1/5–1/4 indehiscent, connective not splitting, sides sparsely to densely lanate, hairs white, to 1 mm, sutures denticulate, teeth to 0.1 mm;

staminode (8–)11–15 mm, included, 0.5–0.1(–1.2) mm diam., tip straight to recurved, distal (2–)4–5 mm sparsely to moderately villous, hairs yellow or white, to 1 mm;

style (11–)13–21 mm.

calyx lobes lanceolate, 5–9 × 0.8–1.9 mm, apex acuminate, sparsely glandular-pubescent;

corolla pale lavender to violet, with faint violet nectar guides, ventricose, 20–35 mm, glandular-pubescent externally, ± white-pubescent internally abaxially, tube 5–7 mm, throat abruptly inflated, 8–11 mm diam., slightly 2-ridged abaxially;

stamens included or longer pair reaching orifice, pollen sacs opposite, navicular, 1.2–1.4 mm, dehiscing completely, connective splitting, sides glabrous, rarely sparsely pubescent, hairs white, to 0.2 mm, sutures papillate;

staminode 11–13 mm, reaching orifice, 0.2–0.3 mm diam., tip straight to slightly recurved, distal 2–5 mm sparsely pubescent, hairs yellowish, to 1 mm;

style 14–16 mm.

Capsules

7–11 × 4–6 mm.

7–8 × 4–5 mm, glabrous.

2n

= 96.

Penstemon strictiformis

Penstemon calycosus

Phenology Flowering May–Jun. Flowering May–Jun.
Habitat Juniper and pinyon-juniper woodlands. Woodlands, meadows, rocky slopes, stream banks.
Elevation 1700–2400 m. (5600–7900 ft.) 10–300 m. (0–1000 ft.)
Distribution
from FNA
AZ; CO; NM; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; CT; DC; GA; IL; IN; KY; MA; MD; ME; MI; MN; MO; NC; NJ; NY; OH; PA; RI; SC; TN; VA; VT; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

Plants of Penstemon strictiformis from southwestern Colorado (Montezuma County) and northwestern New Mexico (San Juan County) closely match the type, with large corollas, broad calyx lobes, and large pollen sacs. Plants from southeastern Utah have relatively smaller corollas, narrower calyx lobes, and smaller pollen sacs.

Penstemon strictiformis often is confused with P. strictus; many specimens from southwestern Colorado are difficult to place with certainty due to morphologic intermediacy.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Penstemon calycosus is concentrated in the Central Lowlands and Interior Low Plateaus; occurrences farther south and east, especially in New England, may be from introductions (F. W. Pennell 1935). The range lies almost entirely within that of P. digitalis, but historic ranges of both P. calycosus and P. digitalis are difficult to delimit due to both past and continuing introduction and spread of both species. Calyx lobe lengths to 12 mm are reported in the literature; those appear to be from fruiting calyces. Penstemon calycosus usually has glabrous anthers, but many specimens, especially from Illinois and Ohio, bear hairs on the adaxial surfaces of the anthers. Some populations from this region exhibit variation in vegetative (leaf size and bract shape and margins) and floral characters (calyx lobe shape and size, corolla color, and anther pubescence), suggesting possible hybridization with P. digitalis, an observation that would be consistent with the findings of A. C. Koelling (1964).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 17, p. 180. FNA vol. 17, p. 197.
Parent taxa Plantaginaceae > Penstemon > subg. Penstemon > sect. Glabri Plantaginaceae > Penstemon > subg. Penstemon > sect. Penstemon
Sibling taxa
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. calycosus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
P. abietinus, P. absarokensis, P. acaulis, P. acuminatus, P. alamosensis, P. albertinus, P. albidus, P. albomarginatus, P. ambiguus, P. ammophilus, P. anguineus, P. angustifolius, P. arenarius, P. arenicola, P. aridus, P. arkansanus, P. attenuatus, P. atwoodii, P. auriberbis, P. australis, P. azureus, P. baccharifolius, P. barbatus, P. barnebyi, P. barrettiae, P. bicolor, P. bleaklyi, P. bracteatus, P. breviculus, P. brevisepalus, P. buckleyi, P. caesius, P. caespitosus, P. calcareus, P. californicus, P. canescens, P. cardinalis, P. cardwellii, P. carnosus, P. caryi, P. centranthifolius, P. cinicola, P. clevelandii, P. clutei, P. cobaea, P. comarrhenus, P. compactus, P. concinnus, P. confertus, P. confusus, P. crandallii, P. cusickii, P. cyananthus, P. cyaneus, P. cyanocaulis, P. cyathophorus, P. dasyphyllus, P. davidsonii, P. deamii, P. deaveri, P. debilis, P. degeneri, P. deustus, P. digitalis, P. diphyllus, P. discolor, P. dissectus, P. distans, P. dolius, P. duchesnensis, P. eatonii, P. elegantulus, P. ellipticus, P. eriantherus, P. euglaucus, P. fendleri, P. filiformis, P. flavescens, P. floribundus, P. floridus, P. flowersii, P. franklinii, P. fremontii, P. fruticiformis, P. fruticosus, P. gairdneri, P. gibbensii, P. glaber, P. glandulosus, P. glaucinus, P. globosus, P. goodrichii, P. gormanii, P. gracilentus, P. gracilis, P. grahamii, P. grandiflorus, P. griffinii, P. grinnellii, P. guadalupensis, P. hallii, P. harbourii, P. harringtonii, P. havardii, P. haydenii, P. heterodoxus, P. heterophyllus, P. hirsutus, P. humilis, P. idahoensis, P. immanifestus, P. incertus, P. inflatus, P. jamesii, P. janishiae, P. kingii, P. kralii, P. labrosus, P. laetus, P. laevigatus, P. laevis, P. lanceolatus, P. laricifolius, P. laxiflorus, P. laxus, P. leiophyllus, P. lemhiensis, P. lentus, P. leonardii, P. linarioides, P. longiflorus, P. lyallii, P. marcusii, P. mensarum, P. metcalfei, P. miser, P. moffatii, P. monoensis, P. montanus, P. moriahensis, P. mucronatus, P. multiflorus, P. murrayanus, P. nanus, P. navajoa, P. neomexicanus, P. neotericus, P. newberryi, P. nitidus, P. nudiflorus, P. oklahomensis, P. oliganthus, P. ophianthus, P. osterhoutii, P. ovatus, P. pachyphyllus, P. pahutensis, P. pallidus, P. palmeri, P. papillatus, P. parryi, P. parvulus, P. parvus, P. patens, P. payettensis, P. paysoniorum, P. peckii, P. penlandii, P. pennellianus, P. perpulcher, P. personatus, P. petiolatus, P. pinifolius, P. pinorum, P. platyphyllus, P. pratensis, P. procerus, P. pruinosus, P. pseudoputus, P. pseudospectabilis, P. pudicus, P. pumilus, P. purpusii, P. putus, P. radicosus, P. rattanii, P. retrorsus, P. rhizomatosus, P. richardsonii, P. roezlii, P. rostriflorus, P. rubicundus, P. rupicola, P. rydbergii, P. saxosorum, P. scapoides, P. scariosus, P. secundiflorus, P. seorsus, P. sepalulus, P. serrulatus, P. smallii, P. spatulatus, P. speciosus, P. spectabilis, P. stenophyllus, P. stephensii, P. strictiformis, P. strictus, P. subglaber, P. subserratus, P. subulatus, P. sudans, P. superbus, P. tenuiflorus, P. tenuis, P. teucrioides, P. thompsoniae, P. thurberi, P. tidestromii, P. tiehmii, P. tracyi, P. triflorus, P. triphyllus, P. tubaeflorus, P. uintahensis, P. utahensis, P. venustus, P. virens, P. virgatus, P. wardii, P. washingtonensis, P. watsonii, P. whippleanus, P. wilcoxii, P. wrightii, P. xylus, P. yampaënsis
Synonyms P. strictus subsp. strictiformis
Name authority Rydberg: Bull. Torrey Bot. Club 31: 642. (1905) — (as Pentstemon) Small: Bull. Torrey Bot. Club 25: 470. (1898) — (as Pentstemon)
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